A
accessory fecundation canal
a slender, usually weakly sclerotized, tube running along dorsal wall of common oviduct from gynatrial sac to point of entrance into common oviduct; divided into basal thickening and fecundation pump.
accessory gland
vermiform gland, q.v.; or, accessory salivary gland, q.v.
accessory male genitalia
in some Schizopteridae, complex dorsoabdominal pregenital structures assumed to perform actual insemination after receiving sperm from primary genitalia.
accessory parempodium
in Leptopodomorpha, e.g., small, secondary parempodia; see also pseudopulvillus.
accessory salivary gland
tubular or vesicular gland associated by a duct with principal salivary gland.
accessory scent glands
small glands associated with primary reservoir of metathoracic scent gland.
aedeagus
that portion of phallus distal to phallobase, including proximal phallotheca (phallosoma) and distal endosoma.
aeropyle
fine pores connected to air spaces in outer and inner meshworks of chorion.
air straps
in Belostomatidae, a pair of straplike appendages derived from abdominal segment 8 used like respiratory siphon in Nepidae to obtain atmospheric air.
anal segment
anal tube, q.v.
anal tube
proctiger, q.v. .
androconia
in males of Scutelleridae and some other Pentatomomorpha, unicellular glands grouped in patches on abdominal venter, with hollow bristlelike androconium set into an alveolus.
androtraumatic insemination
in Phallopirates (Heteroptera: Enicocephalidae), presumed mode of insemination when male can pass sperm to female only after breaking off tip of his own copulatory organ.
annulus
(pl., annuli), a ring encircling an article or segment, as in antennae of some Reduviidae; anal tube, q.v.
anteclypeus
inferior (anterior) portion of clypeus, whenever there is a visible transverse line of demarcation.
antenniferous tubercle
a protuberance of head which bears antenna.
antennal fossa
a groove in which antenna is located or concealed, as in some Phymatinae (Reduviidae).
anteapical claw
see preapical claw.
anterior gonapophyses
first valvulae, q.v.
anterior pronotal lobe
anterior portion of pronotum bearing calli.
anterior ramus
posterior connecting leaf of first valvulae.
anterior valvulae
first valulae, q.v.
apical bulb
spermathecal bulb, q.v.
apterous
completely lacking wings.
arolium
(pl., arolia), bristle- or bladderlike, medial pretarsal structure, originating dorsad of unguitractor and between but isolated from bases of claws (sometimes incorrectly called empodium, parempodium, or rarely puvillus); see empodium and pulvillus; dorsal arolium, q.v.; ventral arolium, q.v.
articulatory apparatus
system of plates and apodemes for suspension of phallus and attachment of its motor muscles, drawn out along phallosoma into ligamentary processes, comprising 1) basal plates attached to suspensory apodemes and ponticulus transversalis and 2) dorsal connectives, ending in capitate processes.
auricle
variously shaped structure on metapleuron of adult Cimicomorpha and Pentatomomorpha, assisting in spreading metathoracic scent-gland products of scent gland from ostiole onto evaporatorium.
auxilia
basipulvillus, q.v.

B
basal apparatus
articulatory apparatus, q.v.
basal foramen
entrance to phallic cavity surrounded by basal plates and ponticulus transversalis, closed or not by a septum.
basal plate
2 major plates of articulatory apparatus; sometimes applied to entire articulatory apparatus.
basiconjunctiva
distal membranous part of phallosoma reaching to, but not including, ejaculatory reservoir.
basipulvillus
basal portion of pulvillus in Pentatomomorpha; see distipulvillus.
Berlese's organ
mesospermalege, q.v., or spermalege, q.v., as a whole.
bothrium
(pl., bothria), pit or tubercle from which a trichobothrium arises.
Brindley's glands
in some adult Reduviidae and Pachynomidae, paired glands located in anteriormost portion of abdomen, with openings situated dorsolaterally just posterior to thoracicoabdominal junction.
buccula
(pl., bucculae), a flange of gena, on each side of basal portion of labium.
bug
a term often loosely used for any insects, but strictly applied to members of suborder Heteroptera; true bug, q.v.
bursa copulatrix
(pl., bursae copulatrices), variously formed structure serving as a vagina, as in Miridae.

C
callar area
middle part of pronotum behind collar and containing calli, and corresponding in size to prothoracic body cavity.
callus
(pl., calli), paired or fused impressions or elevations in anterior part of pronotum behind collar.
capitate processes
mushroomlike ends of dorsal connectives (apodemes) of basal plates, on which are inserted protractor muscles of phallus.
capsid
member of family Miridae.
capsula seminalis
(pl., capsulae seminales), spermathecal bulb, q.v.
cardinate coxae
hinged and elongate hind coxae in pagiopodous taxa.
cephalic gland
maxillary glands, q.v.
cephalic neck
constricted posterior part of head, for most part inserted into prothorax.
Chagas' disease
disease of man and other mammals in South and Central America, Mexico, and Texas, caused by flagellate protozoan Trypanosoma cruzi (Trypanosomatidae) and transmitted by assassin bugs (Reduviidae), esp. Triatoma and Rhodnius
chelicera
(pl., chelicerae), grasping claws of forelegs in some Phymatinae (Reduviidae).
clasper
parameres, q.v.
claval commissure
junction of hemelytra along clavus on midline of body posterior to apex of scutellum and anterior to membrane, developed in most Panheteroptera.
claval furrow
claval suture, q.v.
claval suture
suture of forewing separating clavus from corium.
clavopruina
in Corixidae, a narrow, white frosted area along anteriolateral margin of clavus.
clavus
(pl., clavi), usually parallel-sided and sharply pointed anal area of hemelytron.
claw hairs
setiform microtrichia on outer surface of a claw, e.g., in some Miridae.
claw plate
unguitractor (plate), q.v.
coiled duct
in Gerromorpha, accessory fecundation canal, q.v.
coleopteroid
beetlelike in form, often referring to structure of forewings.
collar
rounded or flattened anterior margin of prothorax.
collum
(pl., colla), collar, q.v., of prothorax.
common oviduct
proximal portion of female genital ducts, between lateral oviducts (whether ectodermal or endodermal) and vagina.
conceptaculum seminis
(pl., conceptacula seminis), in many Cimicoidea, mesodermal organs of sperm storage, being a differentiation of mesodermal oviducts.
conjunctiva
(pl., conjunctivae), intersegmental membrane, q.v.; proximal portion of endosoma of phallus in many Heteroptera.
conjunctival appendages
lobes or processes arising from conjunctiva in expanded aedeagus of many Heteroptera.
connexivum
lateral margin of abdomen, formed by dorsal and ventral laterotergites or laterosternites.
copulatory organ
phallus, q.v.
corial glands
in Plokiophilidae, numerous, large, unicellular glands with low conical openings on dorsal surface of corium.
coriopruina
in Corixidae, a white frosted area between anterior apex of corium and clavopruina.
corium
(pl., coria), in Panheteroptera, proximal coriaceous or otherwise differentiated part of forewing exclusive of clavus and distinct from membrane, often being subdivided into anterior (lateral) exocorium and posterior (mesal) endocorium; see cuneus and embolium.
costal fracture
in many Heteroptera, a short, usually transverse line of weakness or break in costal margin of forewing separating sometimes well-differentiated cuneus from rest of corium.
cotton stainer
species of Heteroptera which cause discoloration of cotton fibers by piercing unripe bolls for their sap, e.g., Dysdercus spp. (Pyrrhocoridae).
cricoid sclerite
in many Pentatomomorpha, a parietal differentiation of endosoma delimiting conjunctiva from vesica.
Cryptocerata
Nepomorpha, q.v.
ctenidium
(pl., ctenidia), in Polyctenidae, comblike rows of flattened spines.
cubital furrow
simple or forked furrow or plica on hind wing posterior to Cu.
cultrate
shaped like a pruning knife.
cuneal incisure
costal fracture, q.v.
cuneus
(pl., cunei), in some Heteroptera, usually triangular posterolateral area of corium demarcated by costal fracture.

D
diadenian type
omphalian or diastomian type of metathoracic scent gland with gland cells concentrated within paired glandular components of system, and with scent reservoir(s) differentiated.
diastomian type
in adult Heteroptera, metathoracic scent gland apparatus opening by paired, widely spaced orifices associated with metacoxal cavities on metapleuron; see omphalian type.
distipulvillus
distal membranous or setiform portion of pulvillus in Pentatomomorpha; see basipulvillus.
dorsal abdominal scent gland (s)
in nymphal Heteroptera, 1--4 paired or unpaired ectodermal abdominal gland(s) with paired on unpaired orifices situated intersegmentally or intrasegmentally, structure and function sometimes persisting into adult stage.
dorsal laterotergite
lateral plate of an abdominal tergum, often subdivided into outer and inner laterotergites.
dorsal paratergite
dorsal laterotergite, q.v.
ductus ejaculatorius
(pl., ducti ejaculatorii), median ectodermal efferent duct proximal to phallus, merging into ductus seminis.
ductus seminis
median ectodermal duct in phallus, from foramen ductus to secondary gonopore, frequently differentiated into d. s. proximalis and d. s. distalis.
ductus spermathecae
canal through which sperm enter spermatheca from vagina or bursa copulatrix.

E
ectospermalege
(pl. ectospermalegia), in some Cimicoidea, external pouchlike ectodermal part of spermalege.
egg cap
a lid joined to body of egg along a line of weakness, which is forced off by hatching embryo.
ejaculatory reservoir
in Pentatomomorpha, Dipsocoromorpha, and Nepidae, complex differentiation of proximal end of ductus seminis in endosoma.
embolar groove
trough-shaped groove of forewing running parallel with costal margin anterior to medial fracture and often delimiting embolium; see medial fracture.
embolium
in forewing of some Heteroptera, broadened submarginal part of corium proximal to costal fracture; exocorium, q.v.
empodium
(pl., empodia), distal extension of unguitractor plate, or, often applied to any unpaired structure arising between claws; see arolium, parempodia, and pulvillus.
endocorium
posterior (mesal in repose) part of corium between exocorium and clavus.
endosoma
(pl., endosomata), distal segment of phallus, free of ligamentary processes and surrounding ductus seminis distalis from ejaculatory reservoir (when present) to secondary gonopore.
epipharyngeal sense organ
anterior (x) and posterior (y) groups of sensilla located in epipharynx, apparently with sensory function related to feeding.
Euheteroptera
that taxon including the Dipsocoromorpha, Gerromorpha, Nepomorpha, Leptopodomorpha, Cimicomorpha, and Pentatomomorpha.
evaporative area
evaporatorium, q.v.
evaporatorium
in most Cimicomorpha and Pentatomomorpha, area of specialized cuticule on metathoracic pleuron associated with, and usually surrounding, orifice and auricle of metathoracic scent glands, possibly functioning in controlled dissemination and evaporation of scent gland products.
eversible gland
in adult Saldidae, a gland located in intersegmental membrane dorsal abdominal laterotergites 7 and 8.
exocorium
that part of corium lying between R or R+M and costal margin; see embolium and cuneus.
expansion skating
in Gerroidea, dispersing of fluid (probably saliva) onto water surface, which lowers surface tension causing insect to move much more rapidly than is otherwise possible.
extragenital insemination
traumatic insemination, q.v.

F
false spiracles
in Nepidae, hydrostatic organs, q.v.
fecundation canal
accessory fecundation canal, q.v.
fecundation pump
in some Gerromorpha, a widened area of accessory fecundation canal provided with a pair of plate like flanges.
first gonocoxae
first valifer, q.v.
first gonocoxopodites
first valifer, q.v.
first ramus
connecting leaf of first valvulae.
first valvulae
the outer blades of the ovipositor.
flagellum
(pl., flagella), elongate, sometimes coiled, distal portion of ductus seminis.
forceps
(pl., forcipes), parameres, q.v.
fossula spongiosa
in many Cimicomorpha, apically on one or more pairs of tibiae, a vesicular hemolymph-filled structure beset with adhesive setae.
frenum
(pl., frena), lateral groove in upper margin of scutellum into which fits or catches channeled locking device on lower edge of clavus.

G
genital atrium
vagina, q.v.
genital capsule
pygophore, q.v..
genital chamber
vagina, q.v..
genital segment
in males abdominal segment 9, in females abdominal segments 8 and 9; pygophore, q.v.
glochis
spur or short vein in hind wing arising distally from Cu.
gonapophysis
(pl. gonapophyses), valvula, q.v.
gonocoxite
valifers, q.v.
gonocoxopodite
valifers, q.v.
gonoforceps
(pl., gonoforcipes), parameres, q.v.
gonopod
parameres, q.v.
gonoporal process
flagellum, q.v.
gonopore
secondary gonopore, q.v.
gonostylus
(pl., gonostyli), paramere(s), q.v.
guide
in some male Enicocephalidae, reduced remnants of external genitalia arising from posteroventral margin of pygophore.
gustatory organ
epipharyngeal sense organ, q.v.
Gymnocerata
a grade group of Heteroptera with freely movable, conspicuous antennae.
gynatrial complex
in Gerromorpha, a term referring to that portion of internal ectodermalia composed of gynatrial sac, spermathecal tube, and accessory fecundation canal; also applied to complex of homologous structures in other Heteroptera.
gynatrial glands
ringed glands, q.v.
gynatrial sac
vaginal pouch, q.v.; bursa copulatrix, q.v.
gynatrium
vagina, q.v.

H
hamus
(pl., hami), spur or short vein, sometimes pointed, projecting into middle cell of hind wing and representing M.
harpago
(pl., harpagones), parameres, q.v.
helicoid process
in some Pentatomomorpha, cricoid sclerite, q.v.
hemelytron
(pl., hemelytra), forewing of Heteroptera, especially in Panheteroptera with distinctly thickened proximal portion and membranous distal portion.
hemocoelic insemination
traumatic insemination, q.v.
hemoglobin
respiratory pigment found in hemolymph of Anisopinae (Notonectidae).
hood
in Tinginae (Tingidae), elevated anterior part of prothorax, often covering head.
humerus
(pl., humeri), posterolateral angle of pronotum; humeral angle, q.v.
hydranapheuxis
in Gerromorpha and Leptopodomorpha, process of deforming meniscus of water surface to allow ascension to adjacent substrate.
hygropetric
pertaining to life on a thin film of water on a rock surface, as in some Gerromorpha.
hydrostatic organs
in Nepidae, 3 pairs of ovoid structures on connexiva of abdominal sterna 3--5 near spiracles but not connected to tracheal system, which function in spatial orientation.
hypandrium
(pl., hypandria), ventro-posterior margin of pygophore (abdominal segment 9) when protruded.
hypocostal lamina
ventrally deflected proximal part of costal margin of forewing.
hypocostal ridge
hypocostal lamina, q.v.
hypocular suture
in Corixidae, short sulcus on either side of head capsule posteroventral to eyes.
hypopygium
(pl., hypopygia), pygophore, q.v.

I
intercalary sclerite
in Gerromorpha, Nepomorpha, and a few other groups of Heteroptera, 2 minute sclerotized plates dorsally between segments 3 and 4 of labium.

J
jugum
(pl., juga), mandibular plate q.v.

K

L
lacerate-flush feeding
in phytophagous Heteroptera, process of lacerating and macerating cells with stylets and then flushing out material with saliva and inbibing it; see sawing-clipping feeding.
laciniate ovipositor
ovipositor with elongate, often laterally compressed blades (valvulae).
laminae
an expanded or platelike region, as body margins of Termitaphididae.
larval organ
in many nymphal Saldidae, an apparently sensory structure, in form of depression, located on abdominal sternum 3 just mesad of spiracle.
lateral oviducts
paired canals leading from ovaries to common oviduct, most frequently mesodermal, but in certain Heteroptera proximally mesodermal and distally ectodermal.
laterosternite
lateral subdivisions of sterna of pregenital abdominal segments (e.g., in some aquatic bugs).
laterotergite
dorsal and ventral laterotergites, q.v.
lima
(pl. limae), stridulitrum, q.v.
lorum
(pl., lora), maxillary plate, q.v.

M
M-chromosome
supernumerary autosomes, occurring most commonly in Lygaeoidea and in some Nepomorpha.
macropterous
with both fore- and hind wings fully developed and functional.
macrotrichia
a relatively large or elongate seta, as on abdomen of some Anthocoridae.
male hooks
parameres, q.v.
mandibular plates
that portion of head laterad of (posterior to) clypeus and dorsad of maxillary plate.
mating swarm
a conspicuous cloud of insects, usually males, dancing or hovering over a marker or in lee of an obstruction, serving to attract solitary members of other sex, e.g., Enicocephalomorpha.
maxillary glands
small paired glands opening near bases of maxillae.
maxillary plate
that portion of head ventral to mandibular plate.
medial fracture
longitudinal furrow delimiting exocorium (or embolium) from endocorium; see embolar groove.
mediotergite
unpaired plate of an abdominal tergum with delimited, paired laterotergites.
mesoscutellum
scutellum, q.v.
micropyle
opening in chorion of egg through which sperm pass during process of fertilization, in most Pentatomomorpha manifested externally by elevated tubular or capitate processes.
membrane
membranous apical portion of hemelytron in Panheteroptera and some other Heteroptera.
mesospermalege
(pl. mesospermalegia), in some Cimicoidea, subintegumental mesodermal portion of spermalege into which spermatozoa are injected; see ectospermalege.
metathoracic scent gland
in Heteroptera, universally occurring adult system of paired or unpaired scent glands with single or paired opening on metasternum with external outflow channels (ostiolar canals) which transmit glandular products to ostiole located on metepisterna.
microtrichia
in Gerromorpha, that portion of body hair layer composed of fine spicules.
mutic
without spines

N
natatorial
natatory, fitted for swimming, being generally applied to swimming legs in aquatic bugs (Nepomorpha).
neck
cephalic neck, q.v.
Neoheteroptera
that taxon including the Gerromorpha, Nepomorpha, Leptopodomorpha, Cimicomorpha, and Pentatomomorpha.
nodal furrow
in Corixidae, costal fracture, q.v.
node
costal fracture, q.v.
nymph
immature form; larva, q.v.

O
ocular seta
in many groups of Heteroptera, usually a pair of setae located in disc of compound eye of early instars, often lost later in development.
odoriferous gland
metathoracic scent gland, q.v., or dorsal abdominal scent gland, q.v.
omphalian type
in adult Heteroptera, metathoracic scent gland apparatus usually with a single (rarely double) opening on metasternum (rarely on abdominal sternum 1), with paired on unpaired internal structures; see diastomian type.
omphalium
prominent metasternal opening of omphalian type of metathoracic scent gland.
operculum
(pl., opercula), egg cap, q.v., as for example in most Cimicomorpha.
organ of Berlese
spermalege, q.v.
ostiolar canal
external outflow pathway of metathoracic scent gland, usually leading from metathoracic venter to metepisternum.
ostiolar groove
ostiolar canal, q.v.
ostiolar peritreme
in many Cimicomorpha and Pentatomomorpha, a calloused area of variable shape, surrounding the the ostiole, and itself often surrounded by the evaporatorium.
ostiole
external opening of metathoracic scent gland, often referring to opening on metepisternum.
ovipositor
organ by which eggs are deposited, formed in Heteroptera by paired first and second valvulae.

P
Pagiopoda
that unnatural assemblage of Heteroptera in which posterior coxae are usually elongate and articulation is a hinge joint; see cardinate and Trochalopoda.
pagiopodous
pertaining to Pagiopoda.
pala
(pl., palae), in Corixidae, tarsus of foreleg modified into a hair-fringed scoop for particle feeding, and in males for attachment to females during mating or sexual display.
palm
palma, in Corixidae, that portion of pala, usually pilose, lying between upper and lower row of palmar setae, sometimes furnished with striduatory pegs.
palmar hairs
in Corixidae, usually a row of long setae on lower margin of pala, and a row of short setae along upper margin of palm.
Panheteroptera
that taxon including Nepomorpha, Leptopodomorpha, Cimicomorpha, and Pentatomomorpha.
paraclypeal lobe
mandibular plate, q.v.
paragenital sinus
in Cimicoidea, external pocket or channel leading to external aperture of ectospermalege.
paragenital system
in many Cimicoidea, various structural differentiations in females correlated with traumatic insemination; see spermalege.
paramere
paired male genital structures independent of phallus, arising post embryologically from exterolateral parts of 2 buds (primary phallic lobes), internomedian parts of which give rise to phallus.
parandrium
(pl., parandria), one of a pair of expansions of external wall of pygophore in lateroventral position, provided with setae but not muscles.
paranota
(sing., paranotum), in certain Tingidae, flattened or lamellate sides of pronotum.
parasternites
laterosternites, q.v.
paratergite
laterotergites, q.v.
parempodia
(sing., parempodium), paired setiform or lamellate processes arising distally from unguitractor plate, between claw bases; Schuh); see arolium, empodium, and pulvillus.
pars intermedialis
spermathecal pump, q.v.
peg plates
in Gerromorpha and Ochteridae, minute circular depressions bordered by a shallow rim and filled with a number of subconical pegs, generally found on head and body, and in some species also on certain leg segments.
Pendergrast's organ
specialized organ found on abdominal venter of some Acanthosomatidae.
penis
(pl., penes), phallus, q.v.
penisfilum
(pl., penisfila), in Saldidae, reel system, q.v.
periadenian type
omphalian or diastomian type of metathoracic scent glands, with gland cells uniformly distributed in paired or unpaired components of system without differentiation of scent reservoir.
peritreme
ostiolar peritreme, q.v.
phallandrium
in Phallopirates (Enicocephalidae), conspicuous bulbous copulatory organ composed largely of novel components including genital plates.
phallobase
articulatory apparatus, q.v.
phallosoma
proximal portion of phallus supported by or incorporating ligamentary process and surrounding ductus seminis proximalis to ejaculatory reservoir (if present), often referred to as phallotheca when sclerotized.
phallotheca
sclerotized proximal part of phallosoma (q.v.), especially in Cimicomorpha and Pentatomomorpha.
phallus
(pl., phalli), intromittent organ, including phallobase, aedeagus and its various processes; see endosoma, phallotheca, phallosoma, vesica.
plastron
in Aphelocheiridae and Cryphocricini (Naucoridae), a physical gill formed by a dense mat of microtrichia on ventral body surface.
plate-shaped ovipositor
ovipositor with shortened valvulae which may be fused, reduced, and dorsoventrally compressed; see laciniate ovipositor.
plectrum
ordinarily moveable portion of stridulatory structure; see stridulitrum.
pleustonic
of or pertaining to air-water interface, e.g., Gerridae are pleustonic.
ponticulus transversalis
large dorsal rodlike transverse superior connection between basal plates in male phallus; see articulatory apparatus and basal plates.
pore-bearing plate
in Hyocephalidae, an ovoid pore-bearing structure on each side of abdominal sternum 3.
posterior gonapophyses
second valvulae.
posterior pronotal lobe
posterior expansion of pronotum overlaying part or more rarely all of mesonotum.
posterior ramus
anterior connecting leaf of second valvulae.
posterior valvulae
second valvulae, q.v.
posterior wall
in female Miridae, a sclerotized, platelike structure lying between the rami of the second valvulae.
postnodal pruina
in Corixidae, a white, frosted area along lateral border of corium posterior to nodal furrow (costal fracture).
preapical claws
in Gerroidea, a condition in which pretarsus is inserted proximal to apex of last tarsal segment.
primary gonopore
distal end of ductus ejaculatorius before entering phallus at level of basal foramen to merge into ductus seminis; see secondary gonopore.
principal salivary gland
major salivary gland of paired salivary system, with 2 or more lobes, always associated with an accessory salivary gland.
processus
gonopori, flagellum, q.v.
proctiger
reduced abdominal segment 10, bearing anus, possibly surrounding invaginated abdominal segment 11
pronotal carina
primarily in Tingidae, main or median carina or keel on pronotum.
prosternal furrow
in most Reduviidae, a cross-striated longitudinal groove in prosternum, by means of which stridulation is caused by rubbing apex of rostrum in it by up and down movements of the head.
pseudarolium
(pl., pseudarolia), in Miridae, pulvillus, q.v.
pseudomicropyle
in eggs of Cimicomorpha, hollow chorionic micropylelike processes used for gas exchange.
pseudoperculum
an egg cap without a distinct sealing bar, and in which eclosion is not result of fluid pressure.
pseudoplacental viviparity
viviparity in which eggs contain little or no yolk and embryo presumably receives nourishment from a pseudoplacenta, e.g., in Polyctenidae.
pseudopulvillus
(pl., pseudopulvilli), in Miridae, paired pretarsal structures arising laterally from unguitractor plate, distinct from parempodia and often superficially resembling pulvilli; see also accessory parempodium.
pseudospermathecae
in Pachynomidae, Reduviidae, and Tingidae which lack functional spermatheca, one or 2 saclike or tubular diverticula arising from vagina or common oviduct, functioning as sperm storage storage organs.
pseudospiracle
in Nepidae, hydrostatic organ, q.v.
pulvillus
(pl. pulivilli), in Miridae, some Anthocoridae, nearly all Pentatomomorpha, bladderlike pretarsal structures arising from ventral or mesal surfaces of claws; see basipulvillus, distipulvillus, arolium, empodium, parempodium, and pseudopulvillus.
pygofer
pygophore, q.v..
pygophore
abdominal segment 9 of male, enclosing the phallus.

Q
R
ramus
(pl., rami), connecting leaves (or arms) of ovipositor valvulae.
raptorial
adapted for seizing prey, e.g., forelegs of many predaceous Heteroptera.
receptaculum seminis
spermatheca, q.v.,
rectal
organ, proctiger, q.v.
reel system
in Saldidae, differentiation of ductus seminis at junction of ductus seminis proximalis and distalis into a coiled tube.
remigium
anterior part of wing chiefly involved in flight; the wing anterior to the claval suture.
respiratory siphon
in Nepidae, paired caudal structures derived from abdominal tergum 8, forming a channel in nymphs and a long tube in adults, which connect with eighth abdominal spiracles, and which serve to replenish subhemelytral airstore in these aquatic insects.
Ribaga's organ
ectospermalege or spermalege, q.v.
ring sclerite
in some Lygaeidae, cricoid sclerite, q.v.; paired or unpaired annular sclerotizations encircling ringed glands of vagina, vaginal pouch, or bursa copulatrix.
ringed glands
in some Heteroptera, paired or unpaired glands, dorsally or ventrally on vagina or on vaginal pouch, or bursa copulatrix, sometimes ringed by annular sclerotizations known as ring sclerites.
rostrum
combined labium and maxillary and mandibular stylets.
rotatory coxae
nearly globose hind coxae with a ball and socket articulation; see Trochalopoda

S
salivary sheath
lipoprotein sheath left in plant tissue, formed from hardened salivary secretions, encasing stylets as they penetrate plant tissue.
sawing-clipping feeding
method of feeding in which stylets are moved back and forth in a straight line; see lacerate-flush feeding.
scent glands
dorsal abdominal scent gland in nymphs (sometimes persisting to adulthood) and several types of scent glands in adults (see metathoracic scent glands, Brindley's glands), producing pheromones, allomones, venoms, and other substances, with often notorious and unpleasant smell for humans.
scent pores
ostioles, q.v.
scent reservoir
paired or unpaired reservoir of metathoracic scent glands; see diadenian type, diastomian type and omphalian type.
sclerotized rings
ring sclerites, q.v.
scrobe
a groove, as in foretibia of Phymatinae (Reduviidae) for reception of tarsus.
scutellum
(pl., scutella), triangular part of mesothorax, generally placed between bases of hemelytra, but in some Pentatomoidea partly or completely overlapping them.
sealing bar
in eggs of Cimicomorpha, a bar joining cap to rest of chorion, consisting of a very thin layer of resistant endochorion and a thick amber layer.
second gonocoxae
second valvifers.
second gonocoxopodites
second valvifers.
second ramus
ramus of second valifer.
second valvifers
valifer arising from abdominal segment 9.
second valvulae
median blades of the ovipositor.
secondary gonopore
opening of ductus seminis at or near apex of phallus.
secondary hypocostal ridge
in adult Heteroptera, a secondary modification of hypocostal lamina.
seminal duct
ductus seminis, q.v.
seminal reservoir
ejaculatory reservoir, q.v.
semiring sclerite
in Colobathristidae, cricoid sclerite, q.v.
sieve pore
peg plates, q.v.
sperm reservoir
ejaculatory reservoir, q.v.
spermalege
(pl., spermalegia), in some Cimicoidea, an organ (organs) on pregenital abdominal segments receiving sperm during traumatic insemination and lacking a direct communication with genital apparatus itself; usually consisting of an external integumental pouch (ectospermalege) and an internal mesodermal part (mesospermalege).
spermatheca
(pl., spermathecae), median, dorsal, unpaired, sclerotized diverticulum of vagina serving as sperm storage receptacle; receptaculum seminis, q.v., vermiform gland, q.v., spermathecal tube, q.v.; see also conceptaculum seminis, pseudospermatheca.
spermathecal bulb
generally bulb-shaped terminal portion of spermatheca serving actual sperm storage function.
spermathecal duct
ductus spermathecae, q.v.
spermathecal gland
vermiform gland, q.v.
spermathecal pump
part of spermatheca between ductus spermathecae and spermathecal bulb, frequently differentiated---in true spermatheca---into a muscular pump with flanges.
spermathecal tube
in infraorder Gerromorpha and some other Heteroptera, an elongate, looped, spermatheca with glandular cells in its walls; see spermatheca.
spermatic duct
in many Cimicoidea, duct arising from fusion of vasa deferentia.
spermatic furrow
in many Cimicoidea, groove of left paramere in which runs interlocked phallus.
spermodes
in Cimicidae, intraepithelial network of canals in walls of pedicels and paired oviducts through which spermatozoa pass from conceptacula seminis to ovarioles.
spiracular line
a line drawn through the spiracles on the abdominal venter of trichophoran Pentatomomorpha, used to refer to postion of abdominal trichobothria relative to spiracles.
spongy fossa
fossula spongiosa, q.v.
stapes
basal plates, q.v.
staphylinoid
condition in which hemelytra are reduced and truncate.
static sense organ
hydrostatic organ, q.v.
strainer
in Hyocephalidae, pore-bearing plate, q.v.
stridulitrum
ordinarily stationary portion of stridulatory mechanism; see plectrum.
strigil
strigile, strigilis, stridulitrum, q.v.
stylus
(pl., styli), parameres, q.v.
subgenital plate
in most female Heteroptera abdominal sternum 7, in Enicocephalidae sternum 8.
submacropterous
condition of wings in which corium and clavus of forewings are fully developed with membrane being slightly to greatly reduced, hind wings generally being functional; see brachypterous.
supradistal plate
in some male Enicocephalidae, dorsal cover of genitalia.
suspensorial apodemes
apodemes to which are attached basal plates of articulatory apparatus.
suspensory arms
suspensorial apodemes, q.v.
suspensory processes
suspensorial apodemes, q.v.
swarming
in Enicocephalidae, aggregating in a mating swarm, q.v.
swimming fan
in some Veliidae, e.g., Rhagovelia (Veliidae), fanlike structure usually formed from a modified ventral arolium, and which aids in swimming on flowing water.
synthlipsis
minimum interocular distance.

T
tegmen
(pl., tegmina), tegminal, a forewing not differentiated into proximal coriaceous and distal membranous part, as for example in Enicocephalomorpha; see hemelytron.
third valvulae
a sheathlike structure of the ovipositor, fused with second valifers, absent in all Pentatomomorpha.
traumatic insemination
in many Cimicoidea and some Nabidae, puncturing body wall or wall of inner genitalia by phallus during mating, and deposition of sperm outside usual reproductive tract.
trichobothrium
(pl., trichobothria), specialized, slender, hairlike, sensory setae arising from and including tubercles or pits (bothria) on many body regions and appendages in Heteroptera; see bothrium.
trichomes
modified hairs present on certain myrmecophilous insects which give off secretions that ants (Hymenoptera: Formicidae) imbibe, e.g., base of abdomen in Holoptilinae (Reduviidae).
Trichophora
those members of Heteroptera with trichobothria on pregenital abdominal sterna, i.e., Pentatomomorpha less Aradidae.
Trochalopoda
that grouping of Heteroptera in which posterior coxae are nearly globose and articulation is a ball and socket joint; see Pagiopoda.
trypanosomiasis
a disease caused by infection with Trypanosoma (Trypanosomatidae), transmitted by Triatominae (Reduviidae); see Chagas' Disease.
tylus
(pl., tyli), distal part of clypeus; anteclypeal region.
tymbal
in some Heteroptera, a sound producing membrane on abdominal segment 1 or segments 1 and 2.
tympanal organ
organ sensitive to vibrations, on mesothorax of some Nepomorpha.

U
unguitractor plate
sclerite lying between bases of claws, with which bases of claws articulate distally, to which retractor tendon is attached proximally, and from which parempodia arise distally.

V
vagina
(pl., vaginae), ectodermal genital ducts distal to common oviduct.
vaginal pouch
variably formed pouch that may bear ringed glands, forming part of gynatrial complex.
valvifers
in female Heteroptera, 4 plates or blades, 2 from abdominal segment 8, 2 from abdominal segment 9, articulating on corresponding paratergites and bearing first and second valvulae, respectively.
valvula
the blades (in laciniate type) of the ovipositor, which in 2 pairs, form the egg laying apparatus, and which proximally attached to the body wall via one (or 2) pairs of corresponding rami.
ventral glands
glands located in metathorax of some Reduviidae, distinct from metathoracic scent glands.
ventral laterotergites
ventrally situated laterotergites, distinct from dorsal laterotergites but generally fused with sternum and usually bearing spiracles.
ventral lobe
in Gerromorpha, buccula, q.v.
ventral paratergites
ventral laterotergites, q.v.
ventral plate
in some male Enicocephalidae, fused genital plates.
ventral spine
in Pentatomoidea, a spinelike projection anteriorly from second or third true abdominal sternum, directed toward head and lying at times between coxae.
vermiform gland
in Cimicomorpha, an organ homologous with spermatheca but without sperm-storing function.
vesica
(pl., vesicae), distal part of an endosoma differentiated to form 2 segments; in Lygaeidae, disticonjunctiva, q.v.

W
water bug
member of infraorder Nepomorpha.
wing coupling mechanism
microtrichia-bearing structure at posteroventral margin of clavus of forewing, grasping leading edge of hind wing during expansion and flexion of wings and during flight.
wing-locking mechanism
in Euheteroptera, modification of costal margin of forewing and mesothorax to retain wing firmly in postion in repose (see frenum); see wing-coupling mechanism.

X
x organ
epipharyngeal sense organ, q.v.
xyphus
xiphus, a spinous triangular process of prosternum and/or mesosternum.

Y
y organ
epipharyngeal sense organ, q.v.

Z