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Laurent, R. F. 1997. Morphometric approach of the evolution of the tribe Ptychadenini. Van Wyk, J. H. ed., Proceedings of the FitzSimons Commemorative Symposium (South African lizards: 50 years of progress) and Third H.A.A. Symposium on African herpetology: held at the Transvaal Museum, Pretoria, South Africa, 11–15 October 1993: 183–188. Matieland, Herpetological Association of Africa.