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Brown, J. L., E. Twomey, A. Amézquita, M. B. de Souza, J. P. Caldwell, S. Lötters, R. von May, P. R. Melo-Sampaio, D. Mejía-Vargas, P. E. Pérez-Peña, M. Pepper, E. H. Poelman, M. Sanchez-Rodriguez, and K. Summers. 2011. A taxonomic revision of the Neotropical poison frog genus Ranitomeya (Amphibia: Dendrobatidae). Zootaxa 3083: 1–120.