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Blackburn, D. C., D. P. Bickford, A. C. Diesmos, D. T. Iskandar, and R. M. Brown. 2010. An ancient origin for the enigmatic Flat-headed Frogs (Bombinatoridae: Barbourula) from the islands of Southeast Asia. PLoS (Public Library of Science) One 5 (8: e 12090): 1–8.