Amerana draytonii (Baird and Girard, 1852)

Class: Amphibia > Order: Anura > Family: Ranidae > Genus: Amerana > Species: Amerana draytonii

Rana Draytonii Baird and Girard, 1852, Proc. Acad. Nat. Sci. Philadelphia, 6: 174. Syntypes: Not stated; USNM 11497 (6 specimens) according to Cochran, 1961, Bull. U.S. Natl. Mus., 220: 73. Type locality: "San Francisco, California, and on Columbia River"; restricted to "vicinity of San Francisco", California, USA by Schmidt, 1953, Check List N. Am. Amph. Rept., Ed. 6: 85.

Rana Lecontii Baird and Girard, 1853, Proc. Acad. Nat. Sci. Philadelphia, 6: 301. Syntypes: Not stated; USNM 3362 (2 specimens) according to Cochran, 1961, Bull. U.S. Natl. Mus., 220: 74. Type locality: "San Francisco", San Francisco County, California, USA. Synonymy with Rana draytoni by Cope, 1886, Proc. Am. Philos. Soc., 23: 521; Cope, 1889, Bull. U.S. Natl. Mus., 34: 441; Boulenger, 1919, Ann. Mag. Nat. Hist., Ser. 9, 3: 414.

Rana nigricans Hallowell, 1854, Proc. Acad. Nat. Sci. Philadelphia, 7: 96. Syntypes: Not stated; USNM 3366 and 3376 according to Cochran, 1961, Bull. U.S. Natl. Mus., 220: 76. Type locality: "El Paso Creek", Kern County, California, USA. Synonymy (of Rana longipes) with Rana temporaria aurora by Cope, 1883, Proc. Acad. Nat. Sci. Philadelphia, 35: 28. Synonymy with Rana draytoni by Cope, 1889, Bull. U.S. Natl. Mus., 34: 441; Boulenger, 1919, Ann. Mag. Nat. Hist., Ser. 9, 3: 414. Preoccupied by Rana nigricans Agassiz, 1850.

Rana longipes Hallowell, 1859, Rep. Upon Explor Surv. Route Railroad Mississippi–Pacific Ocean, 10 (Part 4, No. 1): 20. Replacement name for Rana nigricans Hallowell, 1854.

Rana lecontei — Brocchi, 1881, Miss. Scient. Mex. Amer. Centr., Rech. Zool., 3(2, livr. 1): 14; Cochran, 1961, Bull. U.S. Natl. Mus., 220: 74. Incorrect subsequent spelling.

Epirhexis longipes — Yarrow, 1882, Bull. U.S. Natl. Mus., 24: 179.

Rana draytoni draytoni — Cope, 1889, Bull. U.S. Natl. Mus., 34: 441.

Rana aurora draytoni — Camp, 1917, Univ. California Publ. Zool., 17: 115–125.

Rana (Aurorana) draytonii — Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 322.

Rana (Aurorana) draytonii — Shaffer, Fellers, Voss, Oliver, and Pauly, 2004, Mol. Ecol., 13: 2667.

Rana (Laurasiarana, Amerana) draytonii — Hillis and Wilcox, 2005, Mol. Phylogenet. Evol., 34: 311. See Dubois, 2006, Mol. Phylogenet. Evol., 42: 317–330, and Hillis, 2007, Mol. Phylogenet. Evol., 42: 331–338, for discussion. Invalid name formulation under the International Code of Zoological Nomenclature (1999) as discussed by Dubois, 2007, Cladistics, 23: 395.

Rana (Amerana) draytonii — Dubois, 2006, C. R. Biol., Paris, 329: 830; Yuan, Zhou, Chen, Poyarkov, Chen, Jang-Liaw, Chou, Matzke, Iizuka, Min, Kuzmin, Zhang, Cannatella, Hillis, and Che, 2016, Syst. Biol., 65: 835.

Aurorana draytonii — Fei, Ye, and Jiang, 2010, Herpetol. Sinica, 12: 37. See comment under Ranidae record.

Rana (Rana) draytonii — Fouquette and Dubois, 2014, Checklist N.A. Amph. Rept.: 394.

Amerana draytonii — Dubois, Ohler, and Pyron, 2021, Megataxa, 5: 424. 

English Names

Long-footed Frog (Rana longipes [no longer recognized]: Cooper, 1873, Proc. California Acad. Sci., 4: 64). 

Drayton's Frog (Cooper, 1873, Proc. California Acad. Sci., 4: 64). 

Leconte's Frog (Rana lecontei [no longer recognized]: Cooper, 1873, Proc. California Acad. Sci., 4: 64). 

California Red-legged Frog (Grinnell and Storer, 1924, Animal Life in the Yosemite: 666; Storer, 1925, Univ. California Publ. Zool., 27: 43; Wright and Wright, 1933, Handb. Frogs Toads U.S. Canada: xi; Schmidt, 1953, Check List N. Am. Amph. Rept., Ed. 6: 85; Conant, Cagle, Goin, Lowe, Neill, Netting, Schmidt, Shaw, Stebbins, and Bogert, 1956, Copeia, 1956: 176; Stebbins, 1966, Field Guide W. North Am. Rept. Amph.: 71; Liner, 1994, Herpetol. Circ., 23: 27; Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 106; Collins, Huheey, Knight, and Smith, 1978, Herpetol. Circ., 7: 12; Collins, 1997, Herpetol. Circ., 25: 13; Crother, Boundy, Campbell, de Queiroz, Frost, Highton, Iverson, Meylan, Reeder, Seidel, Sites, Taggart, Tilley, and Wake, 2001 "2000", Herpetol. Circ., 29: 14; Stebbins, 2003, Field Guide W. Rept. Amph., Ed. 3: 226; Frost, McDiarmid, and Mendelson, 2008, in Crother (ed.), Herpetol. Circ., 37: 11; Liner and Casas-Andreu, 2008, Herpetol. Circ., 38: 22; Collins and Taggart, 2009, Standard Common Curr. Sci. Names N. Am. Amph. Turtles Rept. Crocodil., ed. 6: 9; Frost, McDiarmid, Mendelson, and Green, 2012, in Crother (ed.), Herpetol. Circ., 39: 20).

Distribution

Mendocino County (California, USA) south along the Pacific coast of the USA to the vicinity of Arroyo Santo Domingo in northern Baja California (Mexico), 0–2440 m elevation. Northern San Joaquin Valley and Sierra Nevada of California. Now extinct in most of southern California as well as from the southern Sierra Nevada.

Geographic Occurrence

Natural Resident: Mexico, United States of America, United States of America - California

Comment

See comment under Amerana aurora for comment on taxonomic confusion with that species. Grismer, 2002, Amph. Rept. Baja California: 78–79, provided an account (as Rana aurora) for the Baja California, Mexico, population. Stebbins, 2003, Field Guide W. Rept. Amph., Ed. 3: 226–227, provided a brief account, figure, and map (and who commented on the disappearance of this species (as Rana aurora) from large areas of its former distribution). Pauly, Ron, and Lerum, 2008, J. Herpetol., 42: 668–679, reported on molecular phylogeography. Fellers, 2005, in Lannoo (ed.), Amph. Declines: 552–554, and Dodd, 2013, Frogs U.S. and Canada, 2 : 715–722, provided accounts that summarized relevant literature. Elliot, Gerhardt, and Davidson, 2009, Frogs and Toads of N. Am.: 242–243, provided an account, photos, and advertisement call. Altig and McDiarmid, 2015, Handb. Larval Amph. US and Canada: 241–242, provided an account of larval morphology and biology.  

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