Tomopterna tuberculosa (Boulenger, 1882)

Class: Amphibia > Order: Anura > Family: Pyxicephalidae > Subfamily: Cacosterninae > Genus: Tomopterna > Species: Tomopterna tuberculosa

Pyxicephalus rugosus Günther, 1865 "1864", Proc. Zool. Soc. London, 1864: 479. Syntypes: BMNH (2 specimens), according to XXX. Type locality: "Pungo Andongo", Angola, and "W. Africa"; specimens noted from "W. Africa" by Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 30, were not considered types by him. Preoccupied by Rana rugosa Temminck and Schlegel, 1838.

Rana tuberculosa Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 30. Replacement name for Pyxicephalus rugosus Günther, 1864.

Rana pulchra Boulenger, 1896, Ann. Mag. Nat. Hist., Ser. 6, 18: 468. Holotype: BMNH, by original designation. Type locality: "Lake Tanganyika". Synonymy by Loveridge, 1957, Bull. Mus. Comp. Zool., 117: 345 (with Rana tuberculosa); Schmidt and Inger, 1959, Explor. Parc Natl. Upemba, Miss. G.F. de Witte, 56: 36.

Rana (Tomopterna) signata Ahl, 1927 "1925", Sitzungsber. Ges. Naturforsch. Freunde Berlin, 1925: 43. Syntypes: ZMB (2 specimens), unnumbered in original publication. Type locality: "Huilla", Angola. Secondary homonym of Polypedates signata Günther, 1872 (= Rana signata). Synonymy by Laurent, 1954, Publ. Cult. Companhia Diamantes Angola, 23: 72.

Rana (Tomopterna) cacondana Ahl, 1927 "1925", Sitzungsber. Ges. Naturforsch. Freunde Berlin, 1925: 44. Holotype: ZMB, unnumbered in original publication. Type locality: "Caconda", Angola. Synonymy by Laurent, 1954, Publ. Cult. Companhia Diamantes Angola, 23: 72.

Rana (Tomopterna) pulchraDe Witte, 1930, Rev. Zool. Bot. Afr., 19: 263.

Pyxicephalus tuberculosusParker, 1936, Novit. Zool., 40: 142. Poynton, 1964, Ann. Natal Mus., 17: 100.

Rana (Tomopterna) tuberculosaMonard, 1937 "1936", Bull. Soc. Neuchatel. Sci. Nat., 62: 44.

Tomopterna cacondaDeckert, 1938, Sitzungsber. Ges. Naturforsch. Freunde Berlin, 1938: 142.

Tomopterna rugosaLaurent, 1954, Publ. Cult. Companhia Diamantes Angola, 23: 72.

Tomopterna tuberculosaLaurent, 1964, Publ. Cult. Companhia Diamantes Angola, 67: 132. Poynton and Broadley, 1985, Ann. Natal Mus., 27: 130-131; Vences, Glaw, Kosuch, Das, and Veith, 2000, Lourenço and Goodman (eds.), Diversité et Endéémisme à Madagascar: 232.

Rana (Tomopterna) tuberculosaDubois, 1981, Monit. Zool. Ital., N.S., Suppl., 15: 233, by implication.

Tomopterna (Tomopterna) tuberculosaDubois, 1987 "1986", Alytes, 5: 57.

English Names

Warty Frog (Cochran, 1961, Living Amph. World: 158).

Beaded Pyxie (Broadley, 1971, Puku, 6: 113; Lambiris, 1990 "1989", Monogr. Mus. Reg. Sci. Nat. Torino, 10: 89).

Beaded Burrowing Frog (Broadley, 1973, J. Herpetol. Assoc. Afr., 10: 23).

Beaded Dwarf Bullfrog (Broadley, 1974, J. Herpetol. Assoc. Afr., 12: 31).

Angola Bullfrog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 110).

Rough Sand Frog (Channing and Howell, 2006, Amph. E. Afr.: 333).

Tuberculate Sand Frog (Channing and Howell, 2006, Amph. E. Afr.: 333 [alternative name]).

Bearded Pyxie (Channing and Howell, 2006, Amph. E. Afr.: 333 [alternative name]).

Bearded Sand Frog (Du Preez and Carruthers, 2009, Compl. Guide Frogs S. Afr.: 448).

Distribution

Angola and northwestern Namibia to Tanzania and Zimbabwe.

Comment

See Laurent, 1964, Publ. Cult. Companhia Diamantes Angola, 67: 133, for synonymy. See accounts by Poynton and Broadley, 1985, Ann. Natal Mus., 27: 130-131, Channing and Howell, 2006, Amph. E. Afr.: 333-334, Pickersgill, 2007, Frog Search: 124-125, and Du Preez and Carruthers, 2009, Compl. Guide Frogs S. Afr.: 448-449. See also (as Rana tuberculosa) Schmidt and Inger, 1959, Explor. Parc Natl. Upemba, Miss. G.F. de Witte, 56: 36-38. Dawood, Channing, and Bogart, 2002, Mol. Phylogenet. Evol., 22: 407-413, reported two unnamed species masquerading under this name from Beira, Mozambique, and Shankara, Namibia. Mercurio, 2011, Amph. Malawi: 296-297, provided an account for Malawi.

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