Tomopterna cryptotis (Boulenger, 1907)

Class: Amphibia > Order: Anura > Family: Pyxicephalidae > Subfamily: Cacosterninae > Genus: Tomopterna > Species: Tomopterna cryptotis

Rana cryptotis Boulenger, 1907, Ann. Mag. Nat. Hist., Ser. 7, 20: 109. Syntypes: BMNH, by original designation, and MCZ 19268 (on exchange from BMNH, according to Barbour and Loveridge, 1946, Bull. Mus. Comp. Zool., 96: 182). Type locality: "Catequero, Ponang Kuma (Dongwenna), and in the Kafitu Swamps", Angola.

Arthroleptis agadezi Angel, 1936, Bull. Soc. Zool. France, 61: 275. Syntypes: MNHNP 36.124-125 (2 specimens), according to the original publication. Type locality: "Agadez (Aïr)", Sahara. Considered a junior synonym of Pyxicephalus delalandii (Tschudi) (Somali specimens of this nominal species) by Angel and Guibé, 1948, Bull. Mus. Natl. Hist. Nat. Paris, Ser. 2, 20: 62, before the firm distinction of Tomopterna cryptotis and Tomopterna delalandii.

Rana delalandii cryptotisLoveridge, 1936, Field Mus. Nat. Hist. Publ., Zool. Ser., 22: 90.

Arthroleptella ahli Deckert, 1938, Sitzungsber. Ges. Naturforsch. Freunde Berlin, 1938: 168. Holotype: ZMB 15, by original designation. Type locality: "Lichtenstein (Süd Afrika)". Synonymy by Poynton, 1964, Ann. Natal Mus., 17: 97.

Tomopterna hieroglyphicaDeckert, 1938, Sitzungsber. Ges. Naturforsch. Freunde Berlin, 1938: 142.

Pyxicephalus delalandii cryptotisPienaar, 1963, Koedoe, 6: 79.

Pyxicephalus delalandei cryptotisPoynton, 1964, Ann. Natal Mus., 17: 96.

Rana (Pyxicephalus) delalandii cryptotisMertens, 1971, Abh. Senckenb. Naturforsch. Ges., 529: 7.

Tomopterna delalandei cryptotisStevens, 1974, Arnoldia, Zimbabwe, 6: 6.

Tomopterna cryptotisPassmore and Carruthers, 1979, S. Afr. Frogs: 120; Poynton and Broadley, 1985, Ann. Natal Mus., 27: 125.

Rana (Tomopterna) cryptotisDubois, 1981, Monit. Zool. Ital., N.S., Suppl., 15: 233, by implication.

Tomopterna (Tomopterna) cryptotisDubois, 1987 "1986", Alytes, 5: 56.

English Names

Striped Pyxie (Pienaar, 1963, Koedoe, 6: 79; Wager, 1965, Frogs S. Afr.: 134; Passmore and Carruthers, 1979, S. Afr. Frogs: 120; Lambiris, 1990 "1989", Monogr. Mus. Reg. Sci. Nat. Torino, 10: 83).

Striped Burrowing Frog (Pienaar, 1963, Koedoe, 6: 79; Passmore and Carruthers, 1979, S. Afr. Frogs: 120).

Common Burrowing Frog (Van Dijk, 1978, J. Herpetol. Assoc. Afr., 17: 15).

Tremolo Sand Frog (Passmore and Carruthers, 1978, J. Herpetol. Assoc. Afr., 19: 5; Passmore and Carruthers, 1979, S. Afr. Frogs: 120; Channing, 2004, in Minter et al. (eds.), Atlas Frogs S. Afr. Lesotho and Swaziland: 321-322).

Striped Sand Frog (Passmore and Carruthers, 1978, J. Herpetol. Assoc. Afr., 19: 5).

Catequero Bullfrog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 110).

Cryptic Sand Frog (Channing and Howell, 2006, Amph. E. Afr.: 329).

Tremolo Sand Frog (Du Preez and Carruthers, 2009, Compl. Guide Frogs S. Afr.: 434).

Common Sand Frog (Largen and Spawls, 2010, Amph. Rept. Ethiopia Eritrea: 194).

Distribution

Provisional: subsaharan Africa in xeric regions north to Senegal, Mauritania, northern Mali, Niger, and Tanzania, southwards to Angola, Namibia, western Lesotho, and Rep. South Africa north of the Western and Eastern Cape Provinces; no records for Uganda or Kenya.

Comment

See Carruthers and Carruthers, 1979, J. Herpetol. Assoc. Afr., 20: 3-5, for distribution records. See Lanza, 1981, Monit. Zool. Ital., N.S., Suppl., 15: 168, and Poynton and Broadley, 1985, Ann. Natal Mus., 27: 125-127, for account and synonymy. See Salvador, 1996, Smithson. Herpetol. Inform. Serv., 109, for account. Rödel, 2000, Herpetofauna W. Afr., 1: 76-78, provided an account. Pickersgill, 2007, Frog Search: 118-121, provided an account and suggested that this nominal species is composed of several lineages. See Largen, 2001, Tropical Zool., 14: 330-331, for comments on distribution. Padial, 2003, Herpetol. Rev., 34: 260, reported the species for Mauritania. Channing and Howell, 2006, Amph. E. Afr.: 329-330, provided an account for East Africa. Padial and De la Riva, 2004, Rev. Esp. Herpetol., 18: 95, provided records from Mauritania. Channing, 2004, in Minter et al. (eds.), Atlas Frogs S. Afr. Lesotho and Swaziland: 321-322, provided an account for South Africa. Du Preez and Carruthers, 2009, Compl. Guide Frogs S. Afr.: 434-435, provided an account for southern Africa. Schiøtz, 1963, Vidensk. Medd. Dansk Naturhist. Foren., 125: 30, provided records for Nigeria (as Tomopterna delalandii). Bates and Haacke, 2003, Navors. Nas. Mus. Bloemfontein, 19: 124-125, discussed the species in Lesotho. See account, photograph, and map for Ethiopia by Largen and Spawls, 2010, Amph. Rept. Ethiopia Eritrea: 194-196. Zimkus and Larson, 2011, Zootaxa, 2933: 27-45, provided molecular data showing that records from Eritrea, Ethiopia, Djibouti, and Somalia belong to other species. The implication is that all populations from north and west of Zaire should have their status reevaluated. Mercurio, 2011, Amph. Malawi: 291, provided an account for Malawi. Channing, Rödel, and Channing, 2012, Tadpoles of Africa: 356–357, provided information on comparative larval morphology.

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