Ptychadena nilotica (Seetzen, 1855)

Class: Amphibia > Order: Anura > Family: Ptychadenidae > Genus: Ptychadena > Species: Ptychadena nilotica

Rana nilotica Seetzen, 1855, Reisen Syrien, 3: 490. Syntypes: 3 specimens, presumably deposted in ZMB. Type locality: "Kahira", (= Cairo), Egypt. 

Ptychadena niloticaDehling and Sinsch, 2013, Zool. Anz., 253

Rana venusta Werner, 1908 "1907", Sitzungsber. Akad. Wiss. Wien, Phys. Math. Naturwiss. Kl., 116: 1892. Syntypes: NHMW (3 specimens), by original designation; NHMW 12235 considered holotype by Häupl, Tiedemann, and Grillitsch, 1994, Kat. Wiss. Samml. Naturhist. Mus. Wien, 9: 33. Type localities: "im Sudan ausschliesslich bei Mongolla", Sudan (3 specimens). Synonymy with Ptychadena mascareniensis by Loveridge, 1936, Field Mus. Nat. Hist. Publ., Zool. Ser., 22: 94; Guibé and Lamotte, 1957, Bull. Inst. Franç. Afr. Noire, Ser. A, 19: 979. Provisional synonymy on the basis of geography (DRF). 

Rana mascareniensis venusta — Loveridge, 1933, Bull. Mus. Comp. Zool., 74: 370; Loveridge, 1936, Field Mus. Nat. Hist. Publ., Zool. Ser., 22: 94; Mertens, 1938, Senckenb. Biol., 20: 425. See comment.

Ptychadena filwoha Largen, 1997, Tropical Zool., 10: 234. Holotype: BMNH 1974.2842, by original designation. Type locality: "Filwoha, Awash National Park, Shoa, Ethiopia (09° 00′ N 39° 58′ E, altitude 800 m)". Provisional synonymy by Zimkus, Lawson, Barej, Barratt, Channing, Dash, Dehling, Du Preez, Gehring, Greenbaum, Gvoždík, Harvey, Kielgast, Kusamba, Nagy, Pabijan, Penner, Rödel, Vences, and Lötters, 2016, Mol. Phylogenet. Evol., 106: 265. 

English Names

Hot Springs Grass Frog (Ptychadena filwoha [no longer recognized]: Largen and Spawls, 2010, Amph. Rept. Ethiopia Eritrea: 176).

Distribution

Nile drainage of Egypt and Sudan into Ethiopia, thence south into eastern Dem. Rep. Congo, Uganda, Rwanda, Burundi, western and southern Kenya and Tanzania, south to northern Botswana, Mozambique, Swaziland, and extreme eastern Rep. South Africa; presumably to be found in the Caprivi Strip of Namibia, Zimbabwe, Zambia, as well as northern Somalia, Djibouti, and southern Eritrea. 

Comment

Resurrected from the synonymy of Ptychadena mascareniensis by Dehling and Sinsch, 2013, Zool. Anz., 253: 143–157, where it had been placed by by Peters, 1869, Reisen Ost-Afr. 1859–1865, 3: 17 (with Rana bibroni); Boulenger, 1879, Bull. Soc. Zool. France, 4: 92-94;Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 52; and Guibé and Lamotte, 1957, Bull. Inst. Franç. Afr. Noire, Ser. A, 19: 978. Formerly noted as Ptychadena cf mascareniensis (Clade A) of Vences, Kosuch, Rödel, Channing, Glaw, and Böhme, 2004, J. Biogeograph., 31: 593–601, who noted another unnamed species in Kenya which they labeled as Ptychadena cf mascareniensis (Clade D). Zimkus, Lawson, Barej, Barratt, Channing, Dash, Dehling, Du Preez, Gehring, Greenbaum, Gvoždík, Harvey, Kielgast, Kusamba, Nagy, Pabijan, Penner, Rödel, Vences, and Lötters, 2016, Mol. Phylogenet. Evol., 106: 254–269, recognized this species as a member o the Ptychadena mascareniensis complex and reported on molecular systematics and range.  Largen and Spawls, 2010, Amph. Rept. Ethiopia Eritrea: 176, provided a brief account for Ptychadena filwoha, and Largen, 2001, Tropical Zool., 14: 337, commented on the range of Ptychadena filwoha

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