Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Phrynomantis annectens Werner, 1910

Class: Amphibia > Order: Anura > Family: Microhylidae > Subfamily: Phrynomerinae > Genus: Phrynomantis > Species: Phrynomantis annectens

Phrynomantis annectens Werner, 1910, Denkschr. Med. Naturwiss. Ges. Jena, 16: 294. Holotype: ZMB 24829, according to Bauer, Günther, and Robeck, 1996, Mitt. Zool. Mus. Berlin, 72: 267. Type locality: "Aar-Rivier", Namibia.

Phrynomantis nasuta Methuen and Hewitt, 1913, Ann. Transvaal Mus., 4: 122. Holotype: TMP 1317, by original designation. Type locality: "a pool at Kraikluft in the Great Karas Mountains at an altitude of 5000 feet", Namibia. The possibility of relationship with Phrynomantis annectans noted by Hewitt, 1926, Ann. S. Afr. Mus., 20: 488. Synonymy by Parker, 1936, Novit. Zool., 40: 143; Loveridge, 1936, Field Mus. Nat. Hist. Publ., Zool. Ser., 22: 111.

Hoplophryne marmorata Ahl, 1934, Zool. Anz., 107: 334. Holotype: ZMB 35132, according to Bauer, Günther, and Robeck, 1996, Mitt. Zool. Mus. Berlin, 72: 263. Type locality: "Okahandja", Namibia. Synonymy by Parker, 1936, Novit. Zool., 40: 143; Loveridge, 1936, Field Mus. Nat. Hist. Publ., Zool. Ser., 22: 111.

Ctenophryne marmorata Ahl, 1935, Zool. Anz., 112: 254. Holotype: ZMB 35377 according to Bauer, Günther, and Robeck, 1996, Mitt. Zool. Mus. Berlin, 72: 261. Type locality: "ohne Fundort und Sammler". Synonymy by Zweifel, 1989, Copeia, 1989: 229-231.

Phrynomerus annectens — Parker, 1936, Novit. Zool., 40: 143.

Phrynomantis annectens — Dubois, 1988, Alytes, 7: 1-5.

Phrynomantis annectans — Bauer and Branch, 2001, Herpetol. Nat. Hist., 8: 126. Incorrect subsequent spelling.

English Names

Marbled Rubber Frog (Passmore and Carruthers, 1978, J. Herpetol. Assoc. Afr., 19: 4; Passmore and Carruthers, 1979, S. Afr. Frogs: 108; Channing, 2001, Amph. Cent. S. Afr.: 232; Du Preez and Carruthers, 2009, Compl. Guide Frogs S. Afr.: 286).

Red-spotted Frog (Van Dijk, 1978, J. Herpetol. Assoc. Afr., 17: 15).

Red-spotted Namibia Frog (Van Dijk, 1978, J. Herpetol. Assoc. Afr., 17: 15).

Red Marbled Frog (Passmore and Carruthers, 1979, S. Afr. Frogs: 108).

Cape Snake-necked Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 91).

Distribution

Namib desert of southwestern Angola south through Namibia to the Augrabies area of western Northern Cape Province, Rep. South Africa.

Geographic Occurrence

Natural Resident: Angola, Namibia, South Africa

Comment

See accounts by Channing, 2001, Amph. Cent. S. Afr.: 232-233, Channing, 2004, in Minter et al. (eds.), Atlas Frogs S. Afr. Lesotho and Swaziland: 198-199, and Du Preez and Carruthers, 2009, Compl. Guide Frogs S. Afr.: 286-287. Bauer and Branch, 2001, Herpetol. Nat. Hist., 8: 126, commented on its range in the Northern Cape Province, South Africa. Channing, Rödel, and Channing, 2012, Tadpoles of Africa: 251–253, provided information on comparative larval morphology. See Ceríaco, de Sá, Bandeira, Valério, Stanley, Kuhn, Marques, Vindum, Blackburn, and Bauer, 2016, Proc. California Acad. Sci., Ser. 4, 63: 15–61, for records of specimens from Namibe Province, southwestern Angola. Marques, Ceríaco, Blackburn, and Bauer, 2018, Proc. California Acad. Sci., Ser. 4, 65 (Suppl. II): 79–80, provided a map for Angola and provided a brief account. Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 108–109, provided a brief account, photograph, and range map. Lobón-Rovira, Vaz Pinto, Becker, Tolley, Measey, Bennet, Boon, de Sá, and Conradie, 2022, Check List, 18: 289–321, reported the species from Iona National Park, southwestern Angola, and characterized its habitat.

External links:

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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.