Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Chiasmocleis leucosticta (Boulenger, 1888)

Class: Amphibia > Order: Anura > Family: Microhylidae > Subfamily: Gastrophryninae > Genus: Chiasmocleis > Species: Chiasmocleis leucosticta

Engystoma leucosticta Boulenger, 1888, Ann. Mag. Nat. Hist., Ser. 6, 1: 416. Holotype: BMNH 1888.4.23.3, according to Parker, 1934, Monogr. Frogs Fam. Microhylidae: 118, now reregistered as 1947.2.11.49 according to museum records. Type locality: "Sierra do Catharina, . . . . Province Santa Catharina", Brazil.

Gastrophryne leucosticta — Stejneger, 1910, Proc. Biol. Soc. Washington, 23: 166.

Nectodactylus spinulosus Miranda-Ribeiro, 1924, Bol. Mus. Nac., Rio de Janeiro, 1: 256. Syntypes: MNRJ 525 (3 specimens) and CAS-SU 115900; MNRJ 525A designated lectotype by Miranda-Ribeiro, 1955, Arq. Mus. Nac., Rio de Janeiro, 42: 409. Type locality: Humboldt—Santa Catharina—Brazil". Given as "Corupá (= Humboldt), Santa Catarina", Brazil by Bokermann, 1966, Lista Anot. Local. Tipo Anf. Brasil.: 77. Synonymy by Carvalho, 1954, Occas. Pap. Mus. Zool. Univ. Michigan, 555: 1-21.

Chiasmocleis leucosticta — Parker, 1934, Monogr. Frogs Fam. Microhylidae: 118.

Chiasmocleis spinulosa — Parker, 1934, Monogr. Frogs Fam. Microhylidae: 117; Dunn, 1949, Am. Mus. Novit., 1419: 6.

Chiasmocleis urbanae Bokermann, 1952, Pap. Avulsos Zool., São Paulo, 10: 277. Holotype: MZUSP 9033, by original designation. Type locality: "Ilha de São Sebastião, Estado de São Paulo, Brasil". Synonymy by Bokermann, 1966, Lista Anot. Local. Tipo Anf. Brasil.: 27, and Cruz, Caramaschi, and Izecksohn, 1997, Alytes, 15: 49-71.

Chiasmocleis (Chiasmocleis) leucosticta — de Sá, Tonini, van Huss, Long, Cuddy, Forlani, Peloso, Zaher, and Haddad, 2018 "2019", Mol. Phylogenet. Evol., 130: 203.

English Names

Santa Catarina Humming Frog (Chiasmocleis leucosticta: Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 88).

Bokermann's Humming Frog (Chiasmocleis urbanae [no longer recognized]: Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 88).

Distribution

States of Santa Catarina and São Paulo, Brazil.

Geographic Occurrence

Natural Resident: Brazil

Endemic: Brazil

Comment

A tetraploid species according to Kasahara and Haddad, 1997, J. Herpetol., 31: 139-142. See account by Cruz, Caramaschi, and Izecksohn, 1997, Alytes, 15: 49-71. In the Chiasmocleis albopunctata clade of Peloso, Sturaro, Forlani, Gaucher, Motta, and Wheeler, 2014, Bull. Am. Mus. Nat. Hist., 386: 20.

External links:

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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.