Cophyla Boettger, 1880

Class: Amphibia > Order: Anura > Family: Microhylidae > Subfamily: Cophylinae > Genus: Cophyla
19 species

Cophyla Boettger, 1880, Zool. Anz., 3: 281. Type species: Cophyla phyllodactyla Boettger, 1880, by monotypy.

Platypelis Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 474. Type species: Platypelis cowanii Boulenger, 1882, by monotypy. Synonymy by Peloso, Frost, Richards, Rodrigues, Donnellan, Matsui, Raxworthy, Biju, Lemmon, Lemmon, and Wheeler, 2016, Cladistics, 32: xxx;  Peloso, Raxworthy, Wheeler, and Frost, 2017, Mol. Phylogenet. Evol., 111: 56–64. 

Platyhyla Boulenger, 1889, Ann. Mag. Nat. Hist., Ser. 6, 4: 247. Type species: Platyhyla grandis Boulenger, 1889, by monotypy. Synonymy by Guibé, 1978, Bonn. Zool. Monogr., 11: 115.

Paracophyla Millot and Guibé, 1951, Mem. Inst. Sci. Madagascar, Ser. A, 5: 209. Type species: Paracophyla tuberculata Millot and Guibé, 1951, by monotypy. Synonymy with Platypelis by Blommers-Schlösser and Blanc, 1991, Faune de Madagascar, 75: 70.

English Names

Whistling Treefrogs (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 89).

Giant Treefrogs (Platypelis [no longer recognized]: Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 91).

Distribution

Madagascar.

Comment

See account by Blommers-Schlösser and Blanc, 1991, Faune de Madagascar, 75: 88–90. Vences, Andreone, and Glaw, 2005, Afr. Zool., 40: 148, noted that the distinctiveness of this genus from Platypelis is arguable. Glaw and Vences, 2007, Field Guide Amph. Rept. Madagascar, Ed. 3: 138–139, provided accounts. Pyron and Wiens, 2011, Mol. Phylogenet. Evol., 61: 543–583, suggested that Cophyla is the sister taxon of one clade of Plethodontohyla (see comment under that generic account). Rakotoarison, Crottini, Müller, Rödel, Glaw, and Vences, 2015, Zootaxa, 3937: 61–89, revised Cophyla and suggested that it is the monophyletic sister taxon of a monophyletic Platypelis. However in the taxon-dense tree (albeit with less data) Platypelis is paraphyletic with respect to Cophyla, and in the tree where the number of Cophyla species is reduced (although with more loci) Platypelis is monophyletic, suggesting that the jury is still out on mutual monophyly of these two genera (DRF). Peloso, Frost, Richards, Rodrigues, Donnellan, Matsui, Raxworthy, Biju, Lemmon, Lemmon, and Wheeler, 2016, Cladistics, 32: xxx, found Platypelis to be paraphyletic with respect to Cophyla and synonymized the genera. See accounts of former Platypelis by Blommers-Schlösser and Blanc, 1991, Faune de Madagascar, 75: 70–87, and Glaw and Vences, 2007, Field Guide Amph. Rept. Madagascar, Ed. 3: 132–137, and who noted that many unnamed species are contained within this genus. Pyron and Wiens, 2011, Mol. Phylogenet. Evol., 61: 543–583, suggested that Platypelis (along with Stumpffia and Rhombophryne) renders Plethodontohyla paraphyletic and they provided a phylogenetic estimate of exemplar species.

Contained taxa (19 sp.):

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