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Copiula Méhely, 1901
Copiula Méhely, 1901, Termés. Füzetek, 24: 193, 242. Type species: Phrynixalus oxyrhinus Boulenger, 1898, by monotypy.
Mehely Frogs (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 89).
See Menzies and Tyler, 1977, J. Zool., London, 183: 431-464, for revision. The systematic status of Copiula, outside of the area covered by Menzies and Tyler, eastern Papua New Guinea, is unresolved according to R. G. Zweifel IN Frost, 1985, Amph. Species World: 367. Burton, 1990, Trans. R. Soc. S. Aust., 114: 87-93, discussed the diagnosis of the genus. Köhler and Günther, 2008, Mol. Phylogenet. Evol., 47: 353-365, suggested on the basis of molecular evidence that Copiula is nonmonophyletic with a group containing Copiula obsti and Copiula pipiens forming the sister taxon of a group containing Austrochaperina derongo, and a group containing Copiula major forming the sister taxon of an unnamed Austrochaperina species. Pyron and Wiens, 2011, Mol. Phylogenet. Evol., 61: 543-583, in their study of Genbank sequences, confirmed this result.
Contained taxa (12 sp.):
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