Boulenophrys spinata (Liu and Hu, 1973)

Class: Amphibia > Order: Anura > Family: Megophryidae > Subfamily: Megophryinae > Genus: Boulenophrys > Species: Boulenophrys spinata

Megophrys spinatus Liu and Hu in Hu, Zhao, and Liu, 1973, Acta Zool. Sinica, 19: 163. Holotype: CIB 63II0615, by original designation. Type locality: "Fang-xiang, Lei-shan Hsien, Kweichow [=Guizhou], altitude 1,100 m", China.

Megophrys (Megophrys) spinataDubois, 1980, Bull. Mens. Soc. Linn. Lyon, 49: 472.

Megophrys (Xenophrys) spinataDubois and Ohler, 1998, Dumerilia, 4: 14; Chen, Zhou, Poyarkov, Stuart, Brown, Lathrop, Wang, Yuan, Jiang, Hou, Chen, Suwannapoom, Nguyen, Duong, Papenfuss, Murphy, Zhang, and Che, 2017, Mol. Phylogenet. Evol., 109: 41.

Xenophrys spinataOhler, 2003, Alytes, 21: 23, by implication; Delorme, Dubois, Grosjean, and Ohler, 2006, Alytes, 24: 17; Chen, Zhou, Poyarkov, Stuart, Brown, Lathrop, Wang, Yuan, Jiang, Hou, Chen, Suwannapoom, Nguyen, Duong, Papenfuss, Murphy, Zhang, and Che, 2017, Mol. Phylogenet. Evol., 106: 41.

Xenophrys (Xenophrys) spinata — Fei and Ye, 2016, Amph. China, 1: 714.  

Megophrys (Panophrys) spinata — Mahony, Foley, Biju, and Teeling, 2017, Mol. Biol. Evol., 34: 755.

Xenophrys (Panophrys) spinata — Chen, Zhou, Poyarkov, Stuart, Brown, Lathrop, Wang, Yuan, Jiang, Hou, Chen, Suwannapoom, Nguyen, Duong, Papenfuss, Murphy, Zhang, and Che, 2017, Mol. Phylogenet. Evol., 109: 41. 

Panophrys spinata — Lyu, Zeng, Wang, Liu, Huang, Li, and Wang, 2021, Zootaxa, 4927: 13. 

Boulenophrys spinata — Dubois, Ohler, and Pyron, 2021, Megataxa, 5: 414; Qi, Lyu, Wang, Mo, Zeng, Zeng, Dai, Li, Grismer, and Wang, 2021, Zootaxa, 5072: 404.

English Names

Spiny Spadefoot Toad (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 86).

Spiny-fingered Horned Toad (Fei, 1999, Atlas Amph. China: 114).

Distribution

Southeastern Sichuan, southern Chongqing, northern and eastern Guizhou, extreme northeastern Yunnan, and northern Guangxi, China, 800 to 1800 m elevation.

Geographic Occurrence

Natural Resident: China, People's Republic of

Endemic: China, People's Republic of

Comment

Related closely to Megophrys omeimontis according to the original publication. Yang, 1991, Amph. Fauna of Yunnan: 62–63; Fei, 1999, Atlas Amph. China: 114–115, provided brief accounts, maps, and figure. Fei and Ye, 2001, Color Handbook Amph. Sichuan: 146, provided a brief account and illustration. Zhang and Wen, 2000, Amph. Guangxi: 48, provided an account (as Megophrys spinatus) for Guangxi. In the Megophrys omeimontis group of Fei, Ye, Huang, Jiang, and Xie, 2005, in Fei et al. (eds.), Illust. Key Chinese Amph.: 79. Yang, 2008, in Yang and Rao (ed.), Amph. Rept. Yunnan: 33–34, provided a brief account for Yunnan, China. Fei, Hu, Ye, and Huang, 2009, Fauna Sinica, Amph. 2: 408–412, provided an account, illustration of the holotype, and spot map. Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 210–211, provided a brief account including photographs. Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 238–239, provided an account, photographs, and a range map. Fei and Ye, 2016, Amph. China, 1: 714–716, provided an account, photographs, and spot map, as Xenophrys spinata. Zhang, 2017, Amph. Rept. Fanjing Mts.: 76–77, provided taxonomic and natural history information for the Fanjing Mountains population in northeastern Guizhou, China. 

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