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Engystomops Jiménez de la Espada, 1872
Engystomops Jiménez de la Espada, 1872, An. Soc. Esp. Hist. Nat., 1: 86. Type species: Engystomops petersi Jiménez de la Espada, 1872, by monotypy.
Microphryne Peters, 1873, Monatsber. Preuss. Akad. Wiss. Berlin, 1873: 616. Type species: Paludicola pustulosa Cope 1864, by monotypy. Coined as a subgenus of Bufo. Synonymy with Engystomops by Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 275; with Eupemphix by Boulenger, 1888, Ann. Mag. Nat. Hist., Ser. 6, 1: 188.
Microphryne — Cope, 1875 "1876", J. Acad. Nat. Sci. Philadelphia, Ser. 2, 8: 155. Treatment as a genus.
Foam Frogs (Liner and Casas-Andreu, 2008, Herpetol. Circ., 38: 12).
Southern Mexico to northern South America south to Bolivia.
Removed from the synonymy of Physalaemus by Nascimento, Caramaschi, and Cruz, 2005, Arq. Mus. Nac., Rio de Janeiro, 63: 312, where it had been placed by Lynch, 1970, Copeia, 1970: 488-496. Previously placed in the synonymy of Eupemphix by Boettger, 1885, Z. Naturwiss., Halle, 58: 436. Ron, Coloma, and Cannatella, 2005, Herpetologica, 61: 178-198, discussed phylogenetics and call structure of the Physalaemus pustulosus group, which largely corresponds to Engystomops. Funk, Angulo, Caldwell, Ryan, and Cannatella, 2008, Herpetologica, 64: 290-304, suggested that the study by Nascimento, Caramaschi, and Cruz, 2005, Arq. Mus. Nac., Rio de Janeiro, 63: 297-320, did not justify the separation of Physalaemus and Engystomops and noted a manuscript in preparation by D.C. Cannatella to substantiate this statement. Pyron and Wiens, 2011, Mol. Phylogenet. Evol., 61: 543-583, in their study of Genbank sequences, confirmed the monophyly of Engystomops, its sister-taxon relationship to a clade composed of Physalaemus and Eupemphix, and provided a tree of exemplar species. Funk, Caminer, and Ron, 2011, Proc. R. Soc. London, Ser. B, Biol. Sci., 279: 1806-1814, suggested that likely five unnamed species exist in Amazonian Ecuador and adjacent Peru, French Guiana, and Amazonian Brazil, based on molecular markers, morphology, and acoustic data.
Contained taxa (9 sp.):
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