Hyla intermedia Boulenger, 1882

Class: Amphibia > Order: Anura > Family: Hylidae > Subfamily: Hylinae > Genus: Hyla > Species: Hyla intermedia

Hylaria variegata Rafinesque, 1814, Specchio Sci., 2, 2: 103. Types: Not designated and not known to exist. Neotype, by subsequent designation of Nascetti, Lanza, and Bullini, 1995, Amphibia-Reptilia, 16: 225, MZUF 20365. Type locality: "comune nei giardini vicino Palermo e Monreale", Sicily, Italy. Neotype from "the wood called Bosco della Ficuzza, about 10 km by air NE of Corleone (Palermo; ca. 37° 00′ N - 14° 20′ E), altitude about 1000 m". Secondary homonym of Rana variegata Bonnaterre, 1789 (= Hyla leucophyllata).

Hyla arborea var. intermedia Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 381. Syntypes: BMNH 52.12.11.55 (Bononia), 82.7.13.6 (Palermo); BMNH 82.7.13.6 designated lectotype by Dubois, 1995, Dumerilia, 2: 67. Type locality: "Bononia" and "Palermo, Sicily", Italy; restricted to "Palermo" by lectotype designation. Synonymy by Dubois, 1995, Dumerilia, 2: 55-71.

Hyla intermediaHerón-Royer, 1890, Bull. Soc. Zool. France, 15: 209.

Hyla italica Nascetti, Lanza, and Bullini, 1995, Amphibia-Reptilia, 16: 225. Replacement name for Hylaria variegata Rafinesque, 1814. Synonymy with Hyla intermedia by Dubois, 1995, Dumerilia, 2: 55-71.

Hyla (Hyla) intermedia — Fouquette and Dubois, 2014, Checklist N.A. Amph. Rept.: 331, by implication. 

English Names

Italian Tree Frog (Arnold, 2002, Rept. Amph. Eur., Ed. 2: 78).

Distribution

Italy north to the southeastern border region of Switzerland and south into Sicily; possibly into Slovenia.

Comment

In the Hyla arborea group of Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005, Bull. Am. Mus. Nat. Hist., 294: 102 (see comment under Hyla). See Dubois, 1995, Dumerilia, 2: 55-71, for extended discussion of nomenclature. Lanza, 1983, Guide Reconoscimento Spec. Animali, Anf. Rett.: 110-113, discussed this species as Hyla arborea. Arnold, 2002, Rept. Amph. Eur., Ed. 2: 74-75, provided a brief account, figure, and map. Canestrelli, Cimmaruta, and Nascetti, 2007, Mol. Ecol., 16: 4808-4821, reported on molecular phylogeography. Stöck, Dubey, Klütsch, Litvinchuk, Scheidt, and Perrin, 2008, Mol. Phylogenet. Evol., 49: 1019-1024, reported on molecular phylogenetics and noted that it is composed of two distinctive subclades. See detailed account for Italy by Lapini, 2007, in Lanza et al. (eds.), Fauna d'Italia, 42 (Amph.): 333-338. See comments by Speybroeck, Beukema, and Crochet, 2010, Zootaxa, 2492: 7, on the evidence of cryptic taxa within this nominal species. Grosse, 2011, Laubfrösche: 1-286, reviewed the biology of the species. Dufresnes, Dubey, Ghali, Canestrelli, and Perrin, 2015, Conserv. Genetics, 16: 1507–1513, reported on introgressive hybridization of an introduced populations with Hyla arborea in western Switzerland. 

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