Dryophytes gratiosus (LeConte, 1856)

Class: Amphibia > Order: Anura > Family: Hylidae > Subfamily: Hylinae > Genus: Dryophytes > Species: Dryophytes gratiosus

Hyla gratiosa Le Conte, 1856, Proc. Acad. Nat. Sci. Philadelphia, 8: 146. Syntypes: Three specimens noted in description, of which ANSP 2089 was recorded as "holotype" by Malnate, 1971, Proc. Acad. Nat. Sci. Philadelphia, 123: 351, this choice substantiated by its formal designation as lectotype by Fouquette and Dubois, 2014, Checklist N.A. Amph. Rept.: 343. Cochran, 1961, Bull. U.S. Natl. Mus., 220: 54, noted USNM 5901 (6 specimens) as syntypes. The status of type(s) is confusing, see Caldwell, 1982, Cat. Am. Amph. Rept., 298: 1, and Fouquette and Dubois, 2014, Checklist N.A. Amph. Rept.: 343, for discussion. Type locality: "lower country of Georgia", considered by Stejneger and Barbour, 1933, Check List N. Am. Amph. Rept., Ed. 3: 35, to be "evidently one of the LeConte plantations in Floyd or Liberty County", Georgia, USA; restricted to "Liberty County, Georgia", USA, by Schmidt, 1953, Check List N. Am. Amph. Rept., Ed. 6: 71, although as not based on disclosed evidence this restriction is invalid according to Fouquette and Dubois, 2014, Checklist N.A. Amph. Rept.: 343.

Epedaphus gratiosusCope, 1885 "1884", Proc. Am. Philos. Soc., 22: 383.

Hyla (Epedaphus) gratiosa — Fouquette and Dubois, 2014, Checklist N.A. Amph. Rept.: 343. 

Dryophytes gratiosus — Duellman, Marion, and Hedges, 2016, Zootaxa, 4104: 23. 

English Names

Florida Hyla (Yarrow, 1882, Bull. U.S. Natl. Mus., 24: 24).

Florida Tree Frog (Dickerson, 1906, The Frog Book: 124; Wright, 1932, Life Hist. Frogs Okefinokee Swamp, 2: 14; Wright and Wright, 1933, Handb. Frogs Toads U.S. Canada: x).

Georgia Tree Frog (Brimley, 1907, J. Elisha Mitchell Sci. Soc., 23: 158).

Bell-frog (Carr, 1940, Univ. Florida Biol. Sci. Ser., 3: 60).

Giant Tree Frog (Wright and Wright, 1933, Handb. Frogs Toads U.S. Canada: x; Viosca, 1949, Pop. Sci. Bull., Louisiana Acad. Sci., 1: 10).

Barking Tree Frog (Schmidt, 1953, Check List N. Am. Amph. Rept., Ed. 6: 71).

Barking Treefrog (Conant, Cagle, Goin, Lowe, Neill, Netting, Schmidt, Shaw, Stebbins, and Bogert, 1956, Copeia, 1956: 176; Conant, 1975, Field Guide Rept. Amph. E. Cent. N. Am., Ed. 2: 321; Collins, Huheey, Knight, and Smith, 1978, Herpetol. Circ., 7: 11; Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 55; Collins, 1997, Herpetol. Circ., 25: 12; Crother, Boundy, Campbell, de Queiroz, Frost, Highton, Iverson, Meylan, Reeder, Seidel, Sites, Taggart, Tilley, and Wake, 2001 "2000", Herpetol. Circ., 29: 10; Frost, McDiarmid, and Mendelson, 2008, in Crother (ed.), Herpetol. Circ., 37: 6; Collins and Taggart, 2009, Standard Common Curr. Sci. Names N. Am. Amph. Turtles Rept. Crocodil., ed. 6: 7; Frost, McDiarmid, Mendelson, and Green, 2012, in Crother (ed.), Herpetol. Circ., 39: 15; Frost, Lemmon, McDiarmid, and Mendelson, 2017, in Crother (ed.), Herpetol. Circ., 43: 12).


Southeastern USA (eastern Virginia to eastern Louisiana); isolated records from southwestern Kentucky, southern New Jersey, Delaware, Tennessee, and northern Alabama.


In the Hyla cinerea group of Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005, Bull. Am. Mus. Nat. Hist., 294: 102. Reviewed by Caldwell, 1982, Cat. Am. Amph. Rept., 298: 1–2. Mitchell, 2005, in Lannoo (ed.), Amph. Declines: 455–456, and Dodd, 2013, Frogs U.S. and Canada, 1: 280–288, provided accounts that summarized the relevant literature. Elliot, Gerhardt, and Davidson, 2009, Frogs and Toads of N. Am.: 48–51, provided an account, photos, and advertisement call. Altig and McDiarmid, 2015, Handb. Larval Amph. US and Canada: 196–198, provided an account of larval morphology and biology. See account of biology and life history in southern Florida by Meshaka and Lane, 2015, Herpetol. Conserv. Biol., 10 (Monogr. 5): 54–56.

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