Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Cochranella granulosa (Taylor, 1949)

Class: Amphibia > Order: Anura > Family: Centrolenidae > Subfamily: Centroleninae > Genus: Cochranella > Species: Cochranella granulosa

Centrolenella granulosa Taylor, 1949, Univ. Kansas Sci. Bull., 33: 265. Holotype: R.C. Taylor 2463, by original designation; now FMNH 178269 according to Marx, 1976, Fieldiana, Zool., 69: 45. Type locality: "Los Diamantes one mile south of Guápiles, [Cantón de Pococí, Provincia Limón,] Costa Rica". Savage, 1974, Rev. Biol. Tropical, 22: 90, commented on the type locality.

Cochranella granulosa — Taylor, 1951, Proc. Biol. Soc. Washington, 64: 34.

Centrolenella granulosa — Savage, 1967, Copeia, 1967: 328.

Cochranella granulosa — Ruiz-Carranza and Lynch, 1991, Lozania, 57: 22.

English Names

Grainy Cochran Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 47).

Granular Glassfrog (Guayasamin, Cisneros-Heredia, McDiarmid, Peña, and Hutter, 2020, Diversity, 12 (222): 78).

Distribution

Humid lowland and premontane slopes from the Atlantic drainage of eastern Honduras to central Panama and on the Pacific versant in humid upland or gallery forests from northern Costa Rica to southwestern Panama, and adjacent Chocó, Colombia, 40–1500 m elevation; seemingly isolated record from El Jardín de los Sueños Private Reserve, Cotopaxi, Ecuador.

Geographic Occurrence

Natural Resident: Colombia, Costa Rica, Ecuador, Honduras, Nicaragua, Panama

Comment

In the Cochranella granulosa group according to Ruiz-Carranza and Lynch, 1991, Lozania, 57: 1-30. Lips and Savage, 1996, Proc. Biol. Soc. Washington, 109: 17–26, included this species (as Cochranella granulosa) in a key to the tadpoles found in Costa Rica. See accounts by Savage, 2002, Amph. Rept. Costa Rica: 363-364, and McCranie and Wilson, 2002, Amph. Honduras: 211-213. McCranie, 2007, Herpetol. Rev., 38: 37, detailed the departmental distribution in Honduras. Kubicki, 2007, Glass Frogs Costa Rica: 140-153, provided an account and detailed range map for Costa Rica. Sunyer, Martínez-Fonseca, Salazar-Saavedra, Galindo-Uribe, and Obando, 2014, Mesoam. Herpetol., 1: 166, provided records for the department of Chontales, Nicaragua. Culebras, Angiolani-Larrea, Tinajero-Romero, Pellet, and Yeager, 2020, Herpetol. Notes, 13: 353–355, noted a record from El Jardín de los Sueños Private Reserve, Cotopaxi, Ecuador. Guayasamin, Cisneros-Heredia, McDiarmid, Peña, and Hutter, 2020, Diversity, 12 (222): 78–81, provided a detailed account (primarily focused on Ecuador), including adult and larval morphology, advertisement call, relationships, natural history, and conservation status. Díaz-Ricaurte and Guevara-Molina, 2022, Stud. Neotrop. Fauna Environ., 57: 51–65, provided a record from Chocó municipality, Chocó Department, northwestern Colombia, and provided notes on natural history. Martínez-Fonseca, Holmes, Sunyer, Westeen, Grundler, Cerda, Fernández-Mena, Loza-Molina, Monagan, Nondorf, Pandelis, and Rabosky, 2024, Check List, 20: 64–65, provided a record from Refugio Bartola, Departamento Río San Juan, southern Nicaragua.

External links:

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For access to general information see Wikipedia
For additional sources of general information from other websites search Google
For access to relevant technical literature search Google Scholar
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For additional information specific to Ecuador see FaunaWebEcuador: Anfibios del Ecuador
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.