Sclerophrys mauritanica (Schlegel, 1841)

Class: Amphibia > Order: Anura > Family: Bufonidae > Genus: Sclerophrys > Species: Sclerophrys mauritanica

Bufo pantherinus Tschudi, 1838, Classif. Batr.: 88. Nomen nudum coined as a synonym of Bufo arabicus Rüppell and Bufo regularis Ruess. Synonymy with Bufo mauritanicus by Boulenger, 1881 "1880", Proc. Zool. Soc. London, 1880: 557, presumably because Bufo pantherinus Duméril and Bibron, 1841, is a synonym of Bufo mauritanicus.

Bufo mauritanicus Schlegel, 1841, in Wagner (ed.), Reisen Regentschaft Algier, 3: 134. Holotype: RMNH 2122, considered the holotype by M.S. Hoogmoed in Frost, 1985, Amph. Species World: 53. This was considered in error by Beukema and Crochet, 2014, Zootaxa, 3754: 498–500, who noted that several specimens constituted a syntypic series and (p. 499) designated RMNH 2122 as lectotype. Type localities: ". . . in den Umgebungen der Stadt Algier. . . . Sümpfen der Ebene Metidscha; . . . Bona", Algeria. Restricted to "in den Umgebungen der Stadt Algier" (= in the vicinity of the city of Algiers) by the lectotype designation. 

Bufo pantherinus Duméril and Bibron, 1841, Erp. Gen., 8: 687. Types: RMNH by implication. Type locality: "l'occident de l'Asie, ainse que le sud-ouest et le nord de l'Afrique . . . . d'Égypte . . . dans les environs de Bone et d'Alger; enfin nous en avons deux provenant de l'Arabie Pétrée"; restricted by Loveridge, 1957, Bull. Mus. Comp. Zool., 117: 151-362, to Egypt. Junior homonym of Bufo pantherinus Smith, 1828. Named as a synonym of Bufo arabicus Rüppell, Bufo regularis Ruess, and attributed to Boie. Synonymy by Boulenger, 1881 "1880", Proc. Zool. Soc. London, 1880: 563. Includes specimens of Bufo regularis and Bufo mauritanicus according to Boulenger, 1881 "1880", Proc. Zool. Soc. London, 1880: 545-574. Considered a synonym of Bufo regularis by Peters, 1882, Naturwiss. Reise Mossambique, Zool. 3: 178, but it is not known if specimens were examined. See discussion in Poynton and Lambiris, 1998, Afr. J. Herpetol., 47: 3-12 (who noted that Bufo pantherinus Smith, 1828, was likely also included in the hypodigm).

"Bufo" mauritanicusFrost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 363. Excluded from Bufo and unassigned to genus.

Amietophrynus mauritanicusVan Bocxlaer, Biju, Loader, and Bossuyt, 2009, BMC Evol. Biol., 9 (e131): 3.

Sclerophrys mauritanica — Ohler and Dubois, 2016, PeerJ, 4(e1553): 9.

English Names

Pantherine Toad (Gadow, 1901, Amphibia and Reptiles: 184).

Moroccan Toad (Ananjeva, Borkin, Darevsky, and Orlov, 1988, Dict. Amph. Rept. Five Languages: 40; CITES).

Berber Toad (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 42).

Morrish Toad (Geniez, Mateo, Geniez, and Pether, 2004, Amph. Rept. W. Sahara: 35).

Distribution

Mediterranean-climate regions of northern Western Sahara and Morocco, North African Spanish territories of Melilla and Ceuta, northern Algeria, and Tunisia; introduced into the vicinity of Algeciras in southern Spain.

Comment

In the Bufo mauritanicus group of Martin, 1972, in Blair (ed.), Evol. Genus Bufo: 61, and Tandy and Keith, 1972, in Blair (ed.), Evol. Genus Bufo: 160. Roussel and Amar, 1985, Alytes, 4: 41-51, detected call differences between Algerian populations that might indicate species distinctiveness. See Salvador, 1996, Smithson. Herpetol. Inform. Serv., 109, for account. Confused with Bufo arabicus Heyden, by Gervais, 1835, Ann. Sci. Nat., Paris, Ser. 2, 6: 312, according to Boulenger, 1881 "1880", Proc. Zool. Soc. London, 1880: 545-574, and Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 298. Lever, 2003, Naturalized Rept. Amph. World: 176, reported on an introduced populationin southern Spain. Geniez, Mateo, Geniez, and Pether, 2004, Amph. Rept. W. Sahara: 35, provided a brief account for Western Sahara. Harris and Perera, 2009, Biologia, Bratislava, 64: 356-360, noted that this species fits with species referred to Amietophrynus; Van Bocxlaer, Biju, Loader, and Bossuyt, 2009, BMC Evol. Biol., 9 (e131): 1-10, independently corroborated this and made the taxonomic change. Beukema, de Pous, Donaire-Barroso, Bogaerts, Garcia-Porta, Escoriza, Arribas, El Mouden, and Carranza, 2013, Zootaxa, 3661: 37, provided an account for Morocco. See comments on ecology and distribution in Marocco (as Bufo mauritanicus) by Guillon, Le Liard, and Slimani, 2004, Bull. Soc. Herpetol.France, 2004: 37–48. 

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