Bufotes boulengeri (Lataste, 1879)

Class: Amphibia > Order: Anura > Family: Bufonidae > Genus: Bufotes > Species: Bufotes boulengeri

Bufo Hemprichi Fitzinger, 1826, Neue Class. Rept.: 65. Type(s): Not stated, presumably originally in NHMW or ZIUW. Type locality: "Ex Africa, Aegypto". Nomen nudum. Synonymy with Bufo viridis by Steindachner, 1867, Reise Österreichischen Fregatte Novara, Zool., Amph.: 40. Placed in this synonymy on the basis of geography (DRF).

Bufo boulengeri  Lataste, 1879, Rev. Internatl. Sci., 2nd year, Paris, 4: 438. Holotype: Lataste collection; now presumably in the BMNH. Type locality: "sur les plateaux de Sersou, entre Tiaret et Frendah, dans la province d'Oran, et à une altitude de 1200 mètres", Algeria.

Bufo viridis var. maculata Camerano, 1884 "1883", Mem. Accad. Sci. Torino, Ser. 2, 35: 233. Types: MZUT. Type locality: "Modica-Sicilia". Presumably a junior synonym of Bufotes balearicus or an available name for the unnamed species of Bufotes boulengeri. Junior homonym of Bufo maculatus Hallowell, 1854.

Pseudepidalea boulengeriFrost, 2007, Amph. Spec. World, vers. 5.0: . by implication of results of Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 365, and Stöck, Moritz, Hickerson, Frynta, Dujsebayeva, Eremchenko, Macey, Papenfuss, and Wake, 2006, Mol. Phylogenet. Evol., 41: 663-689.

Bufo (Bufotes) boulengeriDubois and Bour, 2010, Zootaxa, 2447: 25.

Bufotes boulengeriFrost, 2013, Amph. Spec. World, Vers. 5.6. Required change because of the non-monophyly of Bufo.

English Names

African Green Toad (Beukema, de Pous, Donaire-Barroso, Bogaerts, Garcia-Porta, Escoriza, Arribas, El Mouden, and Carranza, 2013, Zootaxa, 3661: 42). 

Distribution

North Africa from Western Sahara and Morocco east through Algeria, Tunisia, and Libya, to Egypt; island of Lampedusa, Sicily, Italy; populations in western Arabia south from southern Israel and Jordan to Yemen may be assignable to this species (see comment).

Comment

Recognized as a distinct species by Stöck, Moritz, Hickerson, Frynta, Dujsebayeva, Eremchenko, Macey, Papenfuss, and Wake, 2006, Mol. Phylogenet. Evol., 41: 663-689, and removed from the synonymy of Bufo viridis, where it had been placed by Boulenger, 1881 "1880", Proc. Zool. Soc. London, 1880: 553; Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 297. Stöck et al., 2006, also noted an unnamed species in Sicily, Italy, that is the sister taxon of this diploid species. See also Kete, 1992, Turkish J. Zool., 16: 60-70, who seemingly applied the name Bufo arabicus (as a subspecies of Bufo viridis) to this population and who mapped the subspecies in Europe, North Africa, and the Middle East. Balletto, Cherchi, and Gasperetti, 1985, Fauna Saudi Arabia, 7: 343-345, for discussion of populations in Saudi Arabia that may be of this species. See Salvador, 1996, Smithson. Herpetol. Inform. Serv., 109, for account (as Bufo viridis) of North African population. Frynta, Kratochvil, Moravec, Benda, Dandova, Kaftan, Klosova, Mikulova, Nova, and Schwarzova, 2000, Acta Soc. Zool. Bohem., 64: 20, implied that populations (as Bufo viridis) in the southern oases of Libya might require taxonomic recognition. Baha El Din, 2006, Guide Rept. Amph. Egypt: 35-36, provided an account for Egypt (as Bufo viridis). Geniez, Mateo, Geniez, and Pether, 2004, Amph. Rept. W. Sahara: 31-32, provided an account (as Bufo viridis) for Western Sahara. Ibrahim, 2008, Acta Herpetol., Firenze, 3: 35-49, reported the species from southwestern Libya (as Bufo viridis). See comment under Bufotes siculus. Donaire-Barroso, Beukema, de Pous, and del Canto-Gonzalez, 2011, Herpetol. Notes, 3: 71-74, discussed the range in the Rif Mountains of Morocco. Beukema, de Pous, Donaire-Barroso, Bogaerts, Garcia-Porta, Escoriza, Arribas, El Mouden, and Carranza, 2013, Zootaxa, 3661: 42-44, provided a brief account for Morocco. See comments on ecology and distribution in Marocco (as Bufo viridis) by Guillon, Le Liard, and Slimani, 2004, Bull. Soc. Herpetol.France, 2004: 37–48. 

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