Craugastor rugosus (Peters, 1873)

Class: Amphibia > Order: Anura > Superfamily: Brachycephaloidea > Family: Craugastoridae > Subfamily: Craugastorinae > Genus: Craugastor > Species: Craugastor rugosus

Hylodes rugosus Peters, 1873, Monatsber. Preuss. Akad. Wiss. Berlin, 1873: 610. Holotype: ZMB 7812 according to Bauer, Günther, and Klipfel, 1995, in Bauer et al. (eds.), Herpetol. Contr. W.C.H. Peters: 46. Type locality: "Chiriqui", Panama (see comment under Dendrobates maculatus).

Lithodytes pelviculus Cope, 1877, Proc. Am. Philos. Soc., 17: 89. Holotype: USNM 32326 according to Cochran, 1961, Bull. U.S. Natl. Mus., 220: 66. Type locality: "said to have been found on the west coast of Central America, without more specific locality". Synonymy with Hylodes rugosus Peters, 1874, by Noble, 1918, Bull. Am. Mus. Nat. Hist., 38: 329; synonymy with Strabomantis biporcatus Peters, 1863, by Lynch, 1975, Occas. Pap. Mus. Nat. Hist. Univ. Kansas, 38: 1-46.

Hylodes pelviculusBrocchi, 1881, Miss. Scient. Mex. Amer. Centr., Rech. Zool., 3(2, livr. 1): 55; Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 210.

Lithodytes florulentus Cope, 1893, Proc. Am. Philos. Soc., 31: 336. Syntypes: "No. 327 [G.K Cherrie collection]"; now presumed lost according to Savage and Myers, 2002, Am. Mus. Novit., 3357: 38. Type locality: "Boruca", Cantón de Buenos Aires, Provincia de Puntarenas, Costa Rica. Type locality discussed by Savage, 1974, Rev. Biol. Tropical, 22: 80. Synonymy with Eleutherodactylus biporcatus by Lynch, 1975, Occas. Pap. Mus. Nat. Hist. Univ. Kansas, 38: 1-46. Synonymy with Hylodes rugosus Peters, 1873, by Savage and Myers, 2002, Am. Mus. Novit., 3357: 38.

Hylodes florulentusGünther, 1900, Biol. Centr. Amer., Rept. Batr., Vol. 7, Part 157: 234.

Eleutherodactylus pelviculusStejneger, 1904, Annu. Rep. U.S. Natl. Mus. for 1902: 582-583, by implication.

Eleutherodactylus rugosusStejneger, 1904, Annu. Rep. U.S. Natl. Mus. for 1902: 582-583, by implication; Noble, 1918, Bull. Am. Mus. Nat. Hist., 38: 339.

Eleutherodactylus florulentusStejneger, 1904, Annu. Rep. U.S. Natl. Mus. for 1902: 582-583, by implication; Taylor, 1952, Univ. Kansas Sci. Bull., 35: 765.

Eleutherodactylus (Craugastor) florulentusHedges, 1989, in Woods (ed.), Biogeograph. W. Indies: 317, by implication.

Craugastor florulentusCrawford and Smith, 2005, Mol. Phylogenet. Evol., 35: 551, by implication; Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 360.

Craugastor (Craugastor) rugosusHedges, Duellman, and Heinicke, 2008, Zootaxa, 1737: 38.

English Names

Veragua Robber Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 73).

Distribution

Lowlands and premontane slopes of the Pacific versant from the Rio Carara, Puntarenas Province, Costa Rica (9° 47′ N) to extreme western Panama (10–1220 m).

Comment

Removed from the synonymy of Eleutherodactylus biporcatus by Savage and Myers, 2002, Am. Mus. Novit., 3357: 38 (and who provided an account on page 38-39), where it had been placed by Lynch, 1975, Occas. Pap. Mus. Nat. Hist. Univ. Kansas, 38: 1-46. See account by Savage, 2002, Amph. Rept. Costa Rica: 255-256, who included this species in his Eleutherodactylus fitzingeri series, Eleutherodactylus biporcatus group. In the Craugastor gulosus species series of Hedges, Duellman, and Heinicke, 2008, Zootaxa, 1737: 38. In the Craugastor (Craugastorpunctariolus species series of Padial, Grant, and Frost, 2014, Zootaxa, 3825: 123. Köhler, 2011, Amph. Cent. Am.: 140–191, provided a key to the genera and species of Brachycephaloidea (= Craugastoridae, Eleutherodactylidae) in Central America and provided maps and photographs of the species, including this one.

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