Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Leptopelis calcaratus (Boulenger, 1906)

Class: Amphibia > Order: Anura > Family: Arthroleptidae > Subfamily: Leptopelinae > Genus: Leptopelis > Species: Leptopelis calcaratus

Hylambates calcaratus Boulenger, 1906, Ann. Mag. Nat. Hist., Ser. 7, 17: 322. Syntypes: BMNH, by original designation. By museum records these specimens are BMNH 1901.8.1.68 (reregistered 1947.2.10.15), 1901.6.26.27–30 (reregistered 1947.2.10.11–14), 1903.6.20.25–26 (reregisterd 1947.2.10.16–17), 1904.10.26.20 (reregistered 1947.2.10.18), 1901.6.26.27–30 and 1901.8.1.68 (reregistered 1947.2.10.11–15), and 1903.6.20.25–26 (reregistered 1947.2.10.16–17). Type locality: "Efulen", Cameroon, and "Cape St. John and the Rio Benito District (Spanish West Africa [= Equatorial Guinea]".

Leptopelis calcaratus — Noble, 1924, Bull. Am. Mus. Nat. Hist., 49: 234, 237, 332.

Leptopelis (Leptopelis) calcaratus — Laurent, 1941, Rev. Zool. Bot. Afr., 35: 94.

Leptopelis calcaratus calcaratus — Laurent, 1973, Ann. Mus. R. Afr. Cent., Tervuren, Ser. Octavo, Sci. Zool., 202: 50.

Leptopelis calcaratus meridionalis Laurent, 1973, Ann. Mus. R. Afr. Cent., Tervuren, Ser. Octavo, Sci. Zool., 202: 50. Holotype: MRAC 115147, by original designation. Type locality: "Omaniundu, Terr. de Lodja, Sankuru, Zaïre".

English Names

Efulen Forest Treefrog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 69).

Spurred Tree Frog (Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 216).

Distribution

Dense forest from southeastern Nigeria through southern Cameroon to eastern Dem. Rep. Congo, and western Uganda, south through much of Gabon and northern Rep. Congo; Bioko; isolated record from Uigé Province of northwestern Angola.

Geographic Occurrence

Natural Resident: Angola, Cameroon, Central African Republic, Congo, Democratic Republic of the, Congo, Republic of the, Equatorial Guinea, Gabon, Nigeria, Uganda

Comment

De la Riva, 1994, Rev. Esp. Herpetol., 8: 130, provided a record for Equatorial Guinea. Lasso, Rial, Castroviejo, and De la Riva, 2002, Graellsia, 58: 21-34, provided notes on ecological distribution in Equatorial Guinea. Schiøtz, 1963, Vidensk. Medd. Dansk Naturhist. Foren., 125: 87, provided records for Nigeria. Frétey, Barriere, and Ohler, 2006, Herpetol. Rev., 37: 358, provided a record for the Central African Republic and delimited the range. Jackson and Blackburn, 2007, Salamandra, 43: 149-164, provided a record for Nouabale-Ndoki National Park, Rep. Congo. See account for Cameroon by Amiet, 2012, Rainettes Cameroun: 492-498. Channing, Rödel, and Channing, 2012, Tadpoles of Africa: 110, reported on comparative tadpole morphology. Schiøtz, 1999, Treefrogs Afr.: 260–262, provided an account and map. Barej, Pfalzgraff, Hirschfeld, Liedtke, Penner, Gonwouo, Dahmen, Grözinger, Schmitz, and Rödel, 2015, Amph. Rept. Conserv., 9(2, Spec. Section): 56–84, reported on larval morphology. Jongsma, Tobi, Dixon-MacCallum, Bamba-Kaya, Yoga, Mbega, Mve Beh, Emrich, and Blackburn, 2017, Amph. Rept. Conserv., 11 (1: e144): 6, provided a record for southeastern Gabon and briefly discussed habitat and range. Dewynter and Frétey, 2019, Cah. Fondation Biotope, 27: 13, summarized the literature for Gabon. Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 216–217, provided a brief account, photograph, and range map. Mali, Banda, Chifundera, Badjedjea, Sebe, Lokasola, Ewango, Tungaluna, and Akaibe, 2019, Am. J. Zool., 2: 38–43, reported the species from the Okapi Wildlife Reserve, northeastern Dem. Rep. Congo. See brief account, range map, and photographs for Equatorial Guinea in Sánchez-Vialas, Calvo-Revuelta, Castroviejo-Fisher, and De la Riva, 2020, Proc. California Acad. Sci., Ser. 4, 66: 137–230, who suggested that the Bioko I. population might be a different species. Ernst, Lautenschläger, Branquima, and Hölting, 2020, Zoosyst. Evol., 96: 249, noted the species to occur in Uigé Province of northwestern Angola. Greenbaum, Hughes, and Behangana, 2021, Herpetol. Rev., 52: 342, provided a record from Buhoma, Bwindi Impenetrable National Park, Western Region, Uganda, 1523 m. Kako-Wanzalire, Mongo, Ilonga, Mapoli, Mbumba, Neema, Tungaluna, Itoka, and Bogaert, 2021, Tropicultura, 39 (1: 1709): 1–19, briefly discussed habitat preference in north-central Dem. Rep. Congo. Badjedjea, Masudi, Akaibe, and Gvoždík, 2022, Amph. Rept. Conserv., 16 (1: e301): 45, commented on a population from the Kokolopori Bonobo Nature Reserve, Tshuapa Province, Dem. Rep. Congo.

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For access to general information see Wikipedia
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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.