Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Tylototriton ziegleri Nishikawa, Matsui, and Nguyen, 2013

Class: Amphibia > Order: Caudata > Family: Salamandridae > Subfamily: Pleurodelinae > Genus: Tylototriton > Species: Tylototriton ziegleri

Tylototriton ziegleri Nishikawa, Matsui, and Nguyen, 2013, Curr. Herpetol., Kyoto, 32: 43. Holotype: VNMN 3390, by original designation. Type locality: "Mt. Ta Boc, Ban Thang (Thang Village), Tung Vai Municipality, Quan Ba District, Ha Giang Province, Northern Vietnam, (23° 03′ 25″ N, 104° 50′ 47″ E, 1357 m asl)".

Yaotriton ziegleri —Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 87, by implication; Dubois, Ohler, and Pyron, 2021, Megataxa, 5: 408.

Tylototriton (Yaotriton) ziegleri — Raffaëlli, 2013, Urodeles du Monde, 2nd ed.: 187.

English Names

Ziegler's Knobby Newt (Sparreboom, 2014, Salamanders Old World: 382).

Ziegler’s Corcodile [sic] Newt (Poyarkov, Nguyen, Popov, Geissler, Pawangkhanant, Neang, Suwannapoom, and Orlov, 2021, Russ. J. Herpetol., 28 (3A): 68).

Ha Giang Crocodile Newt (Raffaëlli, 2022, Salamanders & Newts of the World: 286).

Distribution

Ha Giang and Cao Bang provinces of northern Vietnam, likely extending into adjacent Lao Cai Province; reported from adjacent Yunnan (Malipo County), China.

Geographic Occurrence

Natural Resident: China, People's Republic of, Vietnam

Comment

The sister taxon of Tylototriton notialis + Tylototriton asperrimus according to the original publication, in which it was noted that this species had been previously confused with Tylototriton vietnamensis and Tylototriton asperrimus. See comment under Tylototriton regarding taxonomy. Raffaëlli, 2013, Urodeles du Monde, 2nd ed.: 187 provided a brief account, photograph, and map. Sparreboom, 2014, Salamanders Old World: 382–384, reviewed the biology, characteristics, distribution, reproduction, and conservation of the species. Bernardes, Le, Nguyen, Pham, Pham, Nguyen, Rödder, Bonkowski, and Ziegler, 2020, ZooKeys, 935: 121–164, provided a range map. Ye, Wei, Han, Ni, Yao, Xu, Li, Rao, and Zhang, 2017, Conserv. Genet. Resources, 9: 503–506, reported on the complete mtDNA genome and reported the species for Malipo County, Yunnan, China. In the Tylototriton (Yaotriton) asperrimus species group of Poyarkov, Nguyen, and Arkhipov, 2021, Taprobanica, 10: 4–22, who discussed phylogenetics. In the Tylototriton asperrimus group of Lyu, Wang, Zeng, Zhou, Qi, Wan, Li, and Wang, 2021, Vert. Zool., Senckenberg, 71: 697–710, who discussed phylogenetics. Liu, Hou, and Rao, 2022, Biodiversity Data J., 10 (e82707): 1–14, discussed the literature and previous records and provided new records from Malipo county, Wenshan Prefecture, Yunnan Province, China. Raffaëlli, 2022, Salamanders & Newts of the World: 286–287, provided an account, summarizing systematics, life history, population status, and distribution (including a polygon map).

External links:

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For access to general information see Wikipedia
For additional sources of general information from other websites search Google
For access to relevant technical literature search Google Scholar
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For additional information specific to China see Amphibia China
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.