Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Isthmura sierraoccidentalis (Lowe, Jones, and Wright, 1968)

Class: Amphibia > Order: Caudata > Family: Plethodontidae > Subfamily: Hemidactyliinae > Genus: Isthmura > Species: Isthmura sierraoccidentalis

Pseudoeurycea belli sierraoccidentalis Lowe, Jones, and Wright, 1968, Contrib. Sci. Nat. Hist. Mus. Los Angeles Co., 140: 1. Holotype: UAZ 12138, by original designation. Type locality: "ca. 11 mi (rd.) E Santa Ana, on old road to Yécora (ca. 0.5 mi. W. Rancho El Puerto), Sonora, Mexico". The locality is estimated to be 21 ± 1 mi west-southwest of Yécora, by old road to Nuri. [This road, connecting Nuri, Sonora, to Bermudez, Chihuahua, and at the time continuing to Yécora, Sonora, is now long abandoned from Bermudez to Yécora and was nearly impassable in 1975 when it was little more than a walking path—DRF.]

Isthmura sierraoccidentalis — Rovito, Parra-Olea, Recuero, and Wake, 2015, Zool. J. Linn. Soc., 175: 185.

English Names

Sonoran False Brook Salamander Liner, 1994, Herpetol. Circ., 23: 13).

Pine-oak Salamander (Liner and Casas-Andreu, 2008, Herpetol. Circ., 38: 32).

Bell's Salamander (Rorabaugh and Lemos-Espinal, 2016, Field Guide Amph. Rept. Sonora: 114).

Distribution

East-central Sonora and adjacent Madrean Chihuahua, Mexico, 750–3300 m elevation.

Geographic Occurrence

Natural Resident: Mexico

Endemic: Mexico

Comment

In the Pseudoeurycea bellii group of Wake and Lynch, 1976, Sci. Bull. Nat. Hist. Mus. Los Angeles Co., 25: 61. Parra-Olea, García-París, Papenfuss, and Wake, 2005, Herpetologica, 61: 145-158, discussed the systematics of the complex. Raffaëlli, 2007, Les Urodèles du Monde: 219-220, provided brief accounts by subspecies, photographs, and map, and considered it to be in the Pseudoeurycea bellii complex of the Pseudoeurycea bellii group. Van Devender, Lowe, and Holm, 1989, Herpetol. Rev., 20: 75, provided a locality in Madrean Chihuahua. Lemos-Espinal, 2007, Anf. Rept. Chihuahua Mexico: 26-27, provided an account for Chihuahua. Raffaëlli, 2013, Urodeles du Monde, 2nd ed.: 285–286, provided brief account, photographs, and map. See comments regarding Pseudoeurycea bellii sierraoccidentalis by Enderson, Van Devender, and Bezy, 2014, Check List, 10: 919. Rorabaugh and Lemos-Espinal, 2016, Field Guide Amph. Rept. Sonora: 114–116, provided a detailed account of natural history, morphology, distribution, and conservation status in Sonora, Mexico. Bryson, Zarza, Grummer, Parra-Olea, Flores-Villela, Klicka, and McCormack, 2018, Mol. Phylogenet. Evol., 125: 75–84, provided a tree and discussed the phylogenomic diversification of the genus across the Mexican highlands. Raffaëlli, 2022, Salamanders & Newts of the World: 642, provided an account summarizing systematics, life history, population status, and distribution (including a polygon map).

External links:

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For access to general information see Wikipedia
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For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.