Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Theloderma bicolor (Bourret, 1937)

Class: Amphibia > Order: Anura > Family: Rhacophoridae > Subfamily: Rhacophorinae > Genus: Theloderma > Species: Theloderma bicolor

Rhacophorus leprosus bicolor Bourret, 1937, Annexe Bull. Gen. Instr. Publique, Hanoi, 1937: 42. Syntypes: (4 specimens) including MNHNP 1938.62, 48.152–153 (formerly LZUH), according to Guibé, 1950 "1948", Cat. Types Amph. Mus. Natl. Hist. Nat.: 52. Type locality: "Chapa" (= Sa Pa), Lao Cai Province, Vietnam.

Theloderma bicolor — Inger, 1985, in Frost (ed.), Amph. Species World: 549; Orlov, Murphy, Ananjeva, Ryabov, and Ho, 2002, Russ. J. Herpetol., 9: 98.

Theloderma (Theloderma) bicolor — Poyarkov, Orlov, Moiseeva, Pawangkhanant, Ruangsuwan, Vassilieva, Galoyan, Nguyen, and Gogoleva, 2015, Russ. J. Herpetol., 22: 276.

English Names

Chapa Bug-eyed Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 114).

Sapa Bug-eyed Frog (Poyarkov, Nguyen, Popov, Geissler, Pawangkhanant, Neang, Suwannapoom, and Orlov, 2021, Russ. J. Herpetol., 28 (3A): 63).

Distribution

Northwestern Vietnam from Lai Chau, Son La, and Lao Cai Provinces south to Quang Tri Province, Vietnam, 1400–1800 m elevation, also reported in Jingdong and Luechun counties, central and southeastern Yunnan, China.

Geographic Occurrence

Natural Resident: China, People's Republic of, Vietnam

Comment

See account (as Rhacophorus leprosus bicolor) by Bourret, 1942, Batr. Indochine: 414–416. Orlov, Murphy, Ananjeva, Ryabov, and Ho, 2002, Russ. J. Herpetol., 9: 98, suggested that this species is very similar to Theoderma corticale and Theloderma kwangsiense. Nguyen, Ho, and Nguyen, 2005, Checklist Amph. Rept. Vietnam: 41, provided specific localities for Vietnam. See Orlov, Dutta, Ghate, and Kent, 2006, Russ. J. Herpetol., 13: 155–175, for brief comments. See photograph, map, description of geographic range and habitat, and conservation status in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 541. Gawor, Chapuis, Pham, Nguyen, Schmitz, and Ziegler, 2012, Zootaxa, 3395: 59–64, reported on larval morphology. Kropachev, Evsyunin, and Orlov, 2013, Russ. J. Herpetol., 20: 56–60, reported on larval morphology. In the subgenus Theloderma, Theloderma leporosum species group of Poyarkov, Orlov, Moiseeva, Pawangkhanant, Ruangsuwan, Vassilieva, Galoyan, Nguyen, and Gogoleva, 2015, Russ. J. Herpetol., 22: 276. Hou, Yu, Chen, Liao, Zhang, Chen, Li, and Orlov, 2017, Russ. J. Herpetol., 24: 99–127, reviewed the species and provided Chinese localities. Reported from Lai Chau Province, Vietnam, by Pham, Hoang, Pham, and Nguyen, 2019, VNU J. Sci. Nat. Sci. Tech., 35: 73–79. Pham, Nguyen, Tran, Le, Nguyen, Nguyen, Pham, Dang, and Doan, 2023, VNU J. Sci. Earth Environ. Sci., 39: 53–61, provided a record for Son La Province, Vietnam.

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For access to general information see Wikipedia
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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For additional information specific to China see Amphibia China
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.