Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Raorchestes signatus (Boulenger, 1882)

Class: Amphibia > Order: Anura > Family: Rhacophoridae > Subfamily: Rhacophorinae > Genus: Raorchestes > Species: Raorchestes signatus

Ixalus signatus Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 106. Syntypes: Including BMNH 1947.2.24.33-34 and 1947.2.27.35-42 according to Bossuyt and Dubois, 2001, Zeylanica, 6: 30, who designated BMNH 1947.2.27.36 as lectotype and described it on page 74-75. Type locality: "Malabar", South India; restricted by Bossuyt and Dubois, 2001, Zeylanica, 6: 30 to "Nilgherries" (= Nilgiri Hillis), Tamil Nadu, India.

Rhacophorus (Philautus) signatus — Ahl, 1931, Das Tierreich, 55: 77.

Philautus signatus — Gorham, 1974, Checklist World Amph.: 166. Inger, Shaffer, Koshy, and Bakde, 1985 "1984", J. Bombay Nat. Hist. Soc., 81: 554.

Philautus (Philautus) signatus — Bossuyt and Dubois, 2001, Zeylanica, 6: 30.

Kirtixalus signatus — Yu, Rao, Zhang, and Yang, 2009, Mol. Phylogenet. Evol., 50: 578.

Pseudophilautus signatus — Li, Che, Murphy, Zhao, Zhao, Rao, and Zhang, 2009, Mol. Phylogenet. Evol., 53: 511.

Raorchestes signatus — Biju, Shouche, Dubois, Dutta, and Bossuyt, 2010, Curr. Sci., Bangalore, 98: 1120, by implication.

English Names

Boulenger's Bubble-nest Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 112).

Quartet Bush Frog (Das and Dutta, 1998, Hamadryad, 23: 67).

Cross-backed Bush Frog (Chanda, 2002, Handb. Indian Amph.: 168; Dinesh, Radhakrishnan, Gururaja, and Bhatta, 2009, Rec. Zool. Surv. India, Occas. Pap., 302: 109).

Cross-backed Nilgiri Bush Frog Dinesh, Radhakrishnan, Deepak, and Kulkarni, 2023, Fauna India Checklist, vers. 5.0 : 14).

Distribution

Nilgiri Hills of the southern Western Ghats, southwestern India (Tamil Nadu: Coonoor, Kothagiri, Udhagamandalam, Naduvattam, Avalanche, and Parsons Valley), below 2000 m elevation, extending into adjacent Kerala.

Geographic Occurrence

Natural Resident: India

Endemic: India

Comment

See accounts by Inger, Shaffer, Koshy, and Bakde, 1985 "1984", J. Bombay Nat. Hist. Soc., 81: 554, and Chanda, 2002, Handb. Indian Amph.: 168. Dutta, 1997, Amph. India Sri Lanka: 86, provided systematic notes, access to literature, and range. See photograph, map, description of geographic range and habitat, and conservation status in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 532. Biju and Bossuyt, 2009, Zool. J. Linn. Soc., 155: 374-444 [431-433], provided an account, distribution map, and discussed relationships. A brief characterization, photograph, and dot map provided by Subramanian, Dinesh, and Radhakrishnan, 2013, Atlas of Endemic Amph. W. Ghats: 154. Sivaprasad, 2013, Common Amph. Kerala: 206–207, provided a brief account, photograph, and dot map for Kerala, India, providing an imprecise record for Kerala.

External links:

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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.