Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Rana tsushimensis Stejneger, 1907

Class: Amphibia > Order: Anura > Family: Ranidae > Genus: Rana > Species: Rana tsushimensis

Rana tsushimensis Stejneger, 1907, Bull. U.S. Natl. Mus., 58: 116. Holotype: USNM 17519 by original designation and Cochran, 1961, Bull. U.S. Natl. Mus., 220: 77. Type locality: "Tsushima, in the Straits of Korea, between the south end of Korea and Kiusiu, Japan".

Rana temporaria tsushimensis — Sato and Takashima, 1955, Misc. Rep. Yamashina Inst. Ornithol. Zool., 6: 262; Kawamura, 1962, J. Sci. Hiroshima Univ., B—Zool., 20: 181-193.

Rana tsushimensis — Kawamura, 1962, J. Sci. Hiroshima Univ., B—Zool., 20: 191; Okada, 1966, Fauna Japon., Anura: 95; Nishioka, Okumoto, Ueda, and Ryuzaki, 1987, Sci. Rep. Lab. Amph. Biol. Hiroshima Univ., 9: 165-212.

Rana (Rana) amurensis tsushimensis — Nakamura and Ueno, 1963, Japan. Rept. Amph. Color: 37.

Rana amurensis tsushimensis — Ueno and Shibata, 1970, Mem. Nat. Sci. Mus. Tokyo, 3: 194.

Rana (Rana) tsushimensis — Dubois, 1987 "1986", Alytes, 5: 41, by implication; Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 333; Yuan, Zhou, Chen, Poyarkov, Chen, Jang-Liaw, Chou, Matzke, Iizuka, Min, Kuzmin, Zhang, Cannatella, Hillis, and Che, 2016, Syst. Biol., 65: 835.

Rana (Laurasiarana) tsishimensis — Hillis and Wilcox, 2005, Mol. Phylogenet. Evol., 34: 311, by implication; see Dubois, 2006, Mol. Phylogenet. Evol., 42: 317-330, and Hillis, 2007, Mol. Phylogenet. Evol., 42: 331-338, for discussion.

English Names

Tsushima Leopard Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 110).

Tsushima Brown Frog (Maeda and Matsui, 1990, Frogs Toads Japan, Ed. 2: 51; Goris and Maeda, 2004, Guide Amph. Rept. Japan: 105).

Distribution

Tsushima I., in the Korea Strait, Japan.

Geographic Occurrence

Natural Resident: Japan

Endemic: Japan

Comment

In the Section Rana, subgenus Rana, Rana temporaria group of Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 333. See Kuramoto, 1974, Copeia, 1974: 815-822, for a discussion of relationships. Reviewed by Okada, 1966, Fauna Japon., Anura: 95-99. Removed from the synonymy of Rana japonica by Kawamura, 1962, J. Sci. Hiroshima Univ., B—Zool., 20: 191, where it had been placed by Boulenger, 1920, Rec. Indian Mus., 20: 93. See Maeda and Matsui, 1989, Frogs Toads Japan, Ed. 1: 51. Most closely related to Rana okinavana according to Tanaka, Matsui, and Takenaka, 1996, Biochem. Syst. Ecol., 24: 302. Maeda and Matsui, 1990, Frogs Toads Japan, Ed. 2: 48-51, provided an account. Goris and Maeda, 2004, Guide Amph. Rept. Japan: 105-107, provided an account, map, and photograph.

External links:

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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.