Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Hylarana papua (Lesson, 1829)

Class: Amphibia > Order: Anura > Family: Ranidae > Genus: Hylarana > Species: Hylarana papua

Rana papua Lesson, 1829, in Duperrey (ed.), Voyage Coquille, Zool., Atlas (Livr. 11): pl. 7, fig. 1 (description by indication). Syntypes: MNHNP 4575 (2 specimens), according to Guibé, 1950 "1948", Cat. Types Amph. Mus. Natl. Hist. Nat.: 42. Clearly including animal figured in the original. Type locality: "Ile de Waigiou" (= Waigeo I.), Papua (New Guinea), Indonesia, acording to Lesson, 1831, in Duperrey (ed.), Zool., Voyage Coquille, 2 (Pt. 1), Livr. 25: 59; rendered as "Offak Bay (now spelt Fak Fak Bay) on the north coast of Waigeo Island" by Menzies, 1992, Sci. New Guinea, 18: 117.

Limnodytes Waigiensis Duméril and Bibron, 1841, Erp. Gen., 6: 514. Types: Originally in MNHNP. Type locality: "l'île Waigiou". Synonymy by Peters and Doria, 1878, Ann. Mus. Civ. Stor. Nat. Genova, 13: 418; Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 64.

Limnodytes papuensis Meyer, 1875 "1874", Monatsber. Preuss. Akad. Wiss. Berlin, 1874: 138. Syntypes: One surviving syntype, MTD 9285, according to Fritz, 2002, Faun. Abh. Staatl. Mus. Tierkd., Dresden, 23: 12, who noted that the remainder of the syntypes were destroyed in WWII. Type locality: "Jobi" Island, West Irian (New Guinea), Indonesia. Synonymy by Peters and Doria, 1878, Ann. Mus. Civ. Stor. Nat. Genova, 13: 418; Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 64.

Limnodytes papua — Peters and Doria, 1878, Ann. Mus. Civ. Stor. Nat. Genova, 13: 418.

Rana papua — Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 188.

Rana (Hylorana) papua — Boulenger, 1918, Ann. Mag. Nat. Hist., Ser. 9, 1: 238. Boulenger, 1920, Rec. Indian Mus., 20: 126, 160.

Rana (Hylarana) papua — Van Kampen, 1923, Amph. Indo-Austral. Arch.: 201. Dubois, 1987 "1986", Alytes, 5: 42; by implication.

Hylorana papua — Deckert, 1938, Sitzungsber. Ges. Naturforsch. Freunde Berlin, 1938: 145.

Rana semelvella Menzies, 1987, Aust. J. Zool., 35: 395. Holotype: UP 2717, by original designation. Type locality: "lily pond in Bulolo township, c. 700 m, Morobe Province, Papua New Guinea". Synonymy by Menzies, 1992, Sci. New Guinea, 18: 115-122. Shea and Kraus, 2007, Zootaxa, 1514: 37-60, commented on types.

Rana (Papurana) semelvella — Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 326.

Rana (Papurana) papua — Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 326.

Sylvirana papua — Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 370.

Hylarana papua — Che, Pang, Zhao, Wu, Zhao, and Zhang, 2007, Mol. Phylogenet. Evol., 43: 3, by implication; Dubois, Ohler, and Pyron, 2021, Megataxa, 5: 423.

Papurana papua — Fei, Ye, and Jiang, 2010, Herpetol. Sinica, 12: 34. See comment under Ranidae record.

Papurana semelvella — Fei, Ye, and Jiang, 2010, Herpetol. Sinica, 12: 34. Undiscussed recognition of taxon. See comment under Ranidae record.

Papurana papua — Oliver, Prendini, Kraus, and Raxworthy, 2015, Mol. Phylogenet. Evol., 90: 187.

English Names

Papua Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 109).

Distribution

Northern lowlands of New Guinea and offshore islands (including Manus and Waigeo), 0–1200 m elevation.

Geographic Occurrence

Natural Resident: Indonesia, Indonesia - Papua Region, Papua New Guinea

Comment

Prior to the revision of Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297, placed in the Section Hylarana, subsection Hydrophylax, subgenus Papurana of Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 326. See comment under Rana florensis. Removed from the synonymy of Rana papua by Inger, 1954, Fieldiana, Zool., 33: 328, where it had been placed by Van Kampen, 1923, Amph. Indo-Austral. Arch.: 201. See accounts by Boulenger, 1920, Rec. Indian Mus., 20: 188, and Menzies, 1987, Aust. J. Zool., 35: 373–418. See Donnellan, Adams, and Aplin, 1989, Herpetologica, 45: 336-343, for taxonomic concerns. Menzies, 2006, Frogs New Guinea & Solomon Is.: 73, provided a brief account. Kraus and Allison, 2007, Herpetol. Monogr., 21: 33–75, provided an account.

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