Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Microhyla malang Matsui, 2011

Class: Amphibia > Order: Anura > Family: Microhylidae > Subfamily: Microhylinae > Genus: Microhyla > Species: Microhyla malang

Microhyla malang Matsui, 2011, Zootaxa, 2814: 39. Holotype: KUHE 17245, by original designation. Type locality: "Gunung Serapi, Matang Range, in the suburbs of Kuching, Kuching Division, Sarawak, Malaysia (01° 36′ N, 110° 11′ E, alt. 290 m a.s.l.; now a part of Kubah National Park)".

English Names

Borneo Rice Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 90, as Microhyla borneensis).

Borneo Narrow-mouthed Frog (Nutphund, 2001, Amph. Thailand: 158; Das, Jankowski, Makmor, and Haas, 2007, Mitt. Hamburg. Zool. Mus. Inst., 104: 156, as Microhyla borneensis).

Bornean Narrow-mouthed Frog (Das, 2007, Amph. Rept. Brunei: 43, as Microhyla borneensis; Haas, Das, Hertwig, Bublies, and Schulz-Schaeffer, 2022, Guide to the Tadpoles of Borneo: 306).

Bornean Chorus Frog (Chan-ard, 2003, Photograph. Guide Amph. Thailand: 102, as Microhyla borneensis).

Long-snouted Frog (Kuehboon-Hee and Sudin, 2008, Herpetol. Rev., 39: 363, as Microhyla borneensis).

Distribution

Western Sarawak and eastern Sabah, Malaysian Borneo, also Brunei and adjacent Kalimantan, Indonesia; the sources of records from Sumatra appears to be based solely on personal communications and require confirmation.

Geographic Occurrence

Natural Resident: Brunei, Indonesia, Malaysia, Malaysia, East (Sarawak and/or Sabah)

Comment

The sister species of Microhyla borneensis, and previously confused with that species, according to the original publication where records of Microhyla borneensis were reidentified as Microhyla malang. Accounts prior to 2011 conflateded Microhyla malang and Microhyla borneensis (= Microhyla nepenthicola and Microhyla malang) as Microhyla borneensis, but presumably are mostly based on Microhyla malang: Inger, 1966, Fieldiana, Zool., 52: 151-154, and Inger and Tan, 1996, Raffles Bull. Zool., 44: 558; Malkmus, Manthey, Vogel, Hoffmann, and Kosuch, 2002, Amph. Rept. Mount Kinabalu: 131-132; Das, Jankowski, Makmor, and Haas, 2007, Mitt. Hamburg. Zool. Mus. Inst., 104: 156; and Kuehboon-Hee and Sudin, 2008, Herpetol. Rev., 39: 363 (who provided a record for Sabah). Firdaus, Ratih, Karima, Kusuma, and Suastika, 2018, Bioinform. Biomed. Res. J., 1: 1–6, reported on the mtDNA phylogenetic relationships of the species of Microhyla within Sumatra, Java, Borneo, and peninsular Malaysia, placing this species as the sister of Microhyla borneensis. In the Microhyla achatina species group of Garg, Suyesh, Das, Jiang, Wijayathilaka, Amarasinghe, Alhadi, Vineeth, Aravind, Senevirathne, Meegaskumbura, and Biju, 2018 "2019", Vert. Zool., Senckenberg, 69: 1–71. Zainudin and Alaudin, 2018, Malaysian Appl. Biol., 47: 15–21, suggested that Microhyla malang is conspecific with Microhyla borneensis on the basis of low genetic distance. Gorin, Solovyeva, Hasan, Okamiya, Karunarathna, Pawangkhanant, de Silva, Juthong, Milto, Nguyen, Suwannapoom, Haas, Bickford, Das, and Poyarkov, 2020, PeerJ, 8 (e9411): 1–47, placed this species in their Microhyla achatina group and suggested that the Sarawak, Sabah, and Kalimantan populations likely represent distinct species. Haas, Das, Hertwig, Bublies, and Schulz-Schaeffer, 2022, Guide to the Tadpoles of Borneo: 306–308, summarized the knowledge of habitat, reproduction, larval morphology and coloration.

External links:

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For access to general information see Wikipedia
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For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.