Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Leptodactylus rhodonotus (Günther, 1869)

Class: Amphibia > Order: Anura > Family: Leptodactylidae > Subfamily: Leptodactylinae > Genus: Leptodactylus > Species: Leptodactylus rhodonotus

Cystignathus rhodonotus Günther, 1869 "1868", Proc. Zool. Soc. London, 1868: 481. Holotype: BMNH 1947.2.17.39 (formerly 1867.6.13.26), according to de Sá, Grant, Camargo, Heyer, Ponssa, and Stanley, 2014, S. Am. J. Herpetol., 9(Spec. Issue 1): 59, and museum records. Type locality: "Chyavetes [=Chayavitas], Eastern Peru".

Gnathophysa rubido Cope, 1874, Proc. Acad. Nat. Sci. Philadelphia, 26: 128. Syntypes: ANSP 11392 and 11394, according to museum records (W.R. Heyer, personal commun.) and MCZ A-4780 (formerly ANSP 11393) according to Barbour and Loveridge, 1929, Bull. Mus. Comp. Zool., 69: 293), which was subsequently designated lectotype by Heyer, 1969, Herpetologica, 25: 3. Type locality: "Moyabamba, [San Martín,] Peru". Synonymy by Heyer, 1969, Herpetologica, 25: 3.

Pleurodema (Gnathophysa) rubida — Knauer, 1878, Naturgesch. Lurche: 106. Incorrect subsequent spelling.

Leptodactylus rubido — Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 243.

Leptodactylus rhodonotus — Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 239.

Leptodactylus rubidus — Boulenger, 1884 "1883", Proc. Zool. Soc. London, 1883: 637. Error for rubido.

English Names

Peru White-lipped Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 82).

Distribution

Andean slopes and the lowlands of the upper Amazon Basin of Bolivia and Peru into westernmost Acre, Brazil, 200–2050 m elevation; reported from Leticia, Colombia.

Geographic Occurrence

Natural Resident: Bolivia, Brazil, Colombia, Peru

Comment

In the Leptodactylus pentadactylus group of Heyer, 1972, Contrib. Sci. Nat. Hist. Mus. Los Angeles Co., 231: 1–8, and Heyer, 1979, Smithson. Contrib. Zool., 301: 1–43. Rodríguez and Duellman, 1994, Univ. Kansas Mus. Nat. Hist. Spec. Publ., 22: 69, provided a brief account for the Iquitos region of northeastern Peru. Köhler, 2000, Bonn. Zool. Monogr., 48: 134–135, provided a brief account. Duellman, 2005, Cusco Amazonico: 289–290, provided an account (adult and larval morphology, description of the call, life history). Angulo, De la Riva, Reichle, and Gascon, 2004, IUCN Red List, e.T57161A11578741: unnumbered, included Leticia, Colombia, and western-most Acre, Brazil, within the distribution but did not cite specimens. Bernarde, Machado, and Turci, 2011, Biota Neotrop., 11: 117–144, reported specimens from Reserva Extrativista Riozinho da Liberdade, Acre, Brazil. In the Leptodactylus pentadactylus species group of de Sá, Grant, Camargo, Heyer, Ponssa, and Stanley, 2014, S. Am. J. Herpetol., 9(Spec. Issue 1): 1–123, and who provided a summary of relevant literature (adult and larval morphology, identification, advertisement call, and range) on pp. 59–60. See brief account for the Manu region, Peru, by Villacampa-Ortega, Serrano-Rojas, and Whitworth, 2017, Amph. Manu Learning Cent.: 236–237. Metcalf, Marsh, Torres Pacaya, Graham, and Gunnels, 2020, Herpetol. Notes, 13: 753–767, reported the species from the Santa Cruz Forest Reserve, Loreto, northeastern Peru. Segalla, Berneck, Canedo, Caramaschi, Cruz, Garcia, Grant, Haddad, Lourenço, Mângia, Mott, Nascimento, Toledo, Werneck, and Langone, 2021, Herpetol. Brasil., 10: 190, included this species as part of the Brazilian fauna.

External links:

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For access to general information see Wikipedia
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For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.