Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Hyperolius guttulatus Günther, 1858

Class: Amphibia > Order: Anura > Family: Hyperoliidae > Subfamily: Hyperoliinae > Genus: Hyperolius > Species: Hyperolius guttulatus

Hyperolius guttulatus Günther, 1858, Arch. Naturgesch., 24: 326. Syntypes: BMNH (3 specimens) according to Günther, 1859 "1858", Cat. Batr. Sal. Coll. Brit. Mus.: 86, these being 1947.2.9.30-31 (formerly 1846.5.5.80) and 1947.2.10.2 (formerly 1848.8.18.25) according to museum records. Type locality: "Afrika".

Hyperolius reticulatus Günther, 1865 "1864", Proc. Zool. Soc. London, 1864: 482. Holotype: BMNH 1947.2.9.56 (formerly1846.5.5.83), by museum records. Type locality: "probably from West Africa", considered as unknown by Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 125. Synonymy by Laurent, 1958, Mem. Inst. Franç. Afr. Noire, 53: 275.

Rappia lagoensis Günther, 1869 "1868", Proc. Zool. Soc. London, 1868: 487. Syntypes: BMNH (2 specimens) by original designation. These being BMNH 1947.2.9.28 (formerly 1865.1.20.18) and 1947.2.9.29 (formerly 1866.3.8.17) according to museum records. Type locality: "Lagos", Nigeria. Given as "Lagos" and "West Africa" by Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 123. Synonymy by Laurent, 1958, Mem. Inst. Franç. Afr. Noire, 53: 275.

Hyperolius lagoensis — Peters, 1875, Monatsber. Preuss. Akad. Wiss. Berlin, 1875: 207; Noble, 1924, Bull. Am. Mus. Nat. Hist., 49: 252, 326; Ahl, 1931, Das Tierreich, 55: 412.

Rappia reticulata — Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 125.

Rappia lagoensis — Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 123.

Rappia guttulata — Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 125.

Hyperolius guttulatus — Noble, 1924, Bull. Am. Mus. Nat. Hist., 49: 252, 326; Ahl, 1931, Das Tierreich, 55: 416.

Hyperolius buchholzi Ahl, 1931, Mitt. Zool. Mus. Berlin, 17: 56 Holotype: ZMB unnumbered, according to the original publication; presumed lost by Tillack, de Ruiter, and Rödel, 2021, Zoosyst. Evol., Berlin, 97: 441. Type locality: "Accra", Ghana. The description in Ahl, 1931, Das Tierreich, 55: 320, appeared subsequently according to Tillack, de Ruiter, and Rödel, 2021, Zoosyst. Evol., Berlin, 97: 441. Synonymy by Schiøtz, 1967, Spolia Zool. Mus. Haun., 25: 161; Rödel, 2000, Herpetofauna W. Afr., 1: 224.

Hyperolius reticulatus — Ahl, 1931, Das Tierreich, 55: 356.

Hyperolius liberiensis Laurent, 1951, Ann. Soc. R. Zool. Belg., 82: 46. Holotype: MNHNP 1921.175, by original designation. Type locality: "Sanikolé = Sanoquelle, Liberia". Synonymy by Laurent, 1958, Mem. Inst. Franç. Afr. Noire, 53: 275.

English Names

Dotted Reed Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 67).

Spotted Reed Frog (Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 188).

Distribution

Sierra Leone to western Gabon in forest; expected in mainland Equatorial Guinea. See comment.

Geographic Occurrence

Natural Resident: Benin, Cameroon, Gabon, Ghana, Guinea, Ivory Coast, Liberia, Nigeria, Sierra Leone, Togo

Likely/Controversially Present: Equatorial Guinea

Comment

Rödel, 2000, Herpetofauna W. Afr., 1: 224–226, provided an account and mentioned misidentifications in the literature. Schiøtz, 1963, Vidensk. Medd. Dansk Naturhist. Foren., 125: 78–80, provided records for Nigeria. Rödel, Brede, Schiefenhövel, Penner, Sinsin, and Nago, 2007, Salamandra, 43: 231–238, reported this species from the Lokoli Forest of southern Benin. Segniagbeto, Bowessidjaou, Dubois, and Ohler, 2007, Alytes, 24: 77, commented on the presence of this species in Togo. See account for Cameroon by Amiet, 2012, Rainettes Cameroun: 350–356. Channing, Rödel, and Channing, 2012, Tadpoles of Africa: 209–210, reported on comparative tadpole morphology. Schiøtz, 1999, Treefrogs Afr.: 120–121, provided a brief account and map. Dewynter and Frétey, 2019, Cah. Fondation Biotope, 27: 25, summarized the literature for Gabon and provided photographs (p. 57). Rödel and Glos, 2019, Zoosyst. Evol., 95: 22, reported this species from the Krahn-Bassa Proposed Protected Area in southeastern Liberia and the Foya Proposed Protected Area in western Liberia. Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 188–189, provided a brief account, photograph, and range map. Portik, Bell, Blackburn, Bauer, Barratt, Branch, Burger, Channing, Colston, Conradie, Dehling, Drewes, Ernst, Greenbaum, Gvoždík, Harvey, Hillers, Hirschfeld, Jongsma, Kielgast, Kouete, Lawson, Leaché, Loader, Lötters, van der Meijden, Menegon, Müller, Nagy, Ofori-Boateng, Ohler, Papenfuss, Rößler, Sinsch, Rödel, Veith, Vindum, Zassi-Boulou, and McGuire, 2019, Syst. Biol., 68: 859–875, implied that this nominal species is a complex. Sánchez-Vialas, Calvo-Revuelta, Castroviejo-Fisher, and De la Riva, 2020, Proc. California Acad. Sci., Ser. 4, 66: 146, considered the species to be a possible member of the Equatorial Guinea fauna. Ernst, Lautenschläger, Branquima, and Hölting, 2020, Zoosyst. Evol., 96: 253, noted the presence of Hyperolius cf. guttulatus in Uigé Province, northwestern Angola. Kanga, Kouamé, Zogbass, Gongomin, Agoh, Kouamé, Konan, Adepo-Gourène, Gourène, and Rödel, 2021, Amph. Rept. Conserv., 15: 71–107, commented on conservation status, identification, range, and habitat on the Ivory Coast side of Mont Nimba. Gansa, Agadjihouèdé, and Hounkanrin, 2023, Afr. Zool., 58: 39–56, reported the species from the lower Ouémé Valley, southern Benin, and briefly descried habitat preference.

External links:

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For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.