Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Osteocephalus verruciger (Werner, 1901)

Class: Amphibia > Order: Anura > Family: Hylidae > Subfamily: Hylinae > Genus: Osteocephalus > Species: Osteocephalus verruciger

Hyla verrucigera Werner, 1901, Verh. Zool. Bot. Ges. Wien, 51: 601. Syntypes: ZMB 16589 and ZMH A00946 according to Hallermann, 1998, Mitt. Hamburg. Zool. Mus. Inst., 95: 198. Type locality: "Ecuador".

Hyla riopastazae Andersson, 1945, Ark. Zool., 37A(2): 72. Holotype: NHRM 1960 according to Duellman, 1977, Das Tierreich, 95: 150. Type locality: "Rio Pastaza, 1840 m", eastern Ecuador; restricted to "Baños, Río Pastaza, Provincia Tungurahua, Ecuador", by Duellman, 1977, Das Tierreich, 95: 150. Synonymy by Trueb and Duellman, 1970, Copeia, 1970: 601.

Hyla orcesi Funkhouser, 1956, Zoologica, New York, 41: 78. Holotype: CAS-SU 13150, by original designation. Type locality: "[Río] Pacayacu, a stream that flows into the Cotapino, drainage of the Suno, Río Napo region, eastern Ecuador; alt. 600-650 mtr." Synonymy by Trueb and Duellman, 1970, Copeia, 1970: 601.

Osteocephalus orcesi — Cochran and Goin, 1970, Bull. U.S. Natl. Mus., 288: 317.

Osteocephalus verrucigerus — Trueb and Duellman, 1970, Copeia, 1970: 601. Incorrect subsequent spelling.

Osteocephalus verruciger — Duellman, 1985, in Frost (ed.), Amph. Species World: 167.

English Names

Ecuador Slender-legged Treefrog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 61).

Distribution

Lower Amazonian slopes of Andes from Colombia (1400–2000 m elevation) to Provincia Morona Santiago, Ecuador, and 203 km to the southeast in the Cerro Kampankis area of Loreto, Peru and into Caqueá, Colombia. See comment about cryptic species.

Geographic Occurrence

Natural Resident: Colombia, Ecuador, Peru

Comment

Almendáriz C. and Orcés, 2004, Rev. Politécnica, Quito, 25: 118, provided distributional data for Ecuador. Ron, Toral, Venegas, and Barnes, 2010, ZooKeys, 70: 67–92, provided an mtDNA which showed samples of Osteocephalus verruciger being noninclusive with respect to Osteocephalus buckleyi, suggesting either introgressive hybridization or cryptic species; they also suggested (p. 85) that records from Peru are based on misidentified Osteocephalus mimeticus. In the Osteocephalus buckleyi species group of Jungfer, Faivovich, Padial, Castroviejo-Fisher, Lyra, Berneck, Iglesias, Kok, MacCulloch, Rodrigues, Verdade, Torres-Gastello, Chaparro, Valdujo, Reichle, Moravec, Gvoždík, Gagliardi-Urrutia, Ernst, De la Riva, Means, Lima, Señaris, Wheeler, and Haddad, 2013, Zool. Scripta, 42: 351–380. Catenazzi and Venegas, 2012, in Pitman et al. (eds.), Rapid Biol. Social Invent. Rep. 24 268, provided the Peruvian record and dicussed the range. Batallas-Revelo and Brito-M., 2016, Rev. Mexicana Biodiversidad, 87: 1292–1300, reported on vocalizations from Sangay National Park, southeastern Ecuador. Chasiluisa, Caminer, Varela-Jaramillo, and Ron, 2020, Neotropical Biodiversity, 6: 21–36, noted that this nominal species is composed of four lineage-species of which the name-bearing clade is found in Napo and Morona-Santiago, Ecuador. Presumably the other species will be named in other papers.

External links:

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For access to general information see Wikipedia
For additional sources of general information from other websites search Google
For access to relevant technical literature search Google Scholar
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For additional information specific to Ecuador see FaunaWebEcuador: Anfibios del Ecuador
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.