Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Nanorana conaensis (Fei and Huang, 1981)

Class: Amphibia > Order: Anura > Family: Dicroglossidae > Subfamily: Dicroglossinae > Genus: Nanorana > Species: Nanorana conaensis

Rana conaensis Fei and Huang In Huang and Fei, 1981, Acta Zootaxon. Sinica, 6: 212. Holotype: NWIPB 770532, by original designation. Type locality: "Mama of Cona Xian, Xizang [= Tibet], altitude 2,900" m, China. Rendered as "Mama, Chuona County, Tibet at the elevation 2900 m" by Huang, Lathrop, and Murphy, 1998, Smithson. Herpetol. Inform. Serv., 118: 6.

Rana (Paa) conaensis — Dubois, 1987 "1986", Alytes, 5: 43.

Paa (Paa) conaensis — Fei, Ye, and Huang, 1990, Key to Chinese Amph.: 157; Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 320.

Rana conaensis — Zhao and Adler, 1993, Herpetol. China: 141.

Nanorana conaensis — Chen, Murphy, Lathrop, Ngo, Orlov, Ho, and Somorjai, 2005, Herpetol. J., 15: 239, by implication; Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 367.

Chaparana (Paa) conaensis — Ohler and Dubois, 2006, Zoosystema, 28: 781.

Nanorana (Paa) conaensis — Che, Zhou, Hu, Papenfuss, Wake, and Zhang, 2010, Proc. Natl. Acad. Sci. USA, Suppl. Inform., doi:10.1073/pnas.1008415107/-/DCSupplemental: 2; Hofmann, Baniya, Litvinchuk, Miehe, Li, and Schmidt, 2019, Ecol. Evol., 9: 14506.

Maculopaa conaensis — Fei, Ye, and Jiang, 2010, Herpetol. Sinica, 12: 26. See comment under Dicroglossidae.

English Names

Cona Paa Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 102).

Cona Spiny Frog (Fei, 1999, Atlas Amph. China: 212; Li, Zhao, and Dong, 2010, Amph. Rept. Tibet: 41).

Cona Hill Frog (Dinesh, Radhakrishnan, Deepak, and Kulkarni, 2023, Fauna India Checklist, vers. 5.0 : 4).

Distribution

Known from the region of the type locality in southern Tibet, China, and in adjacent Bhutan (Haa District); also reported from Khremteng village, Tawang Township, Arunachal Pradesh, India, 2844 m elevation.

Geographic Occurrence

Natural Resident: Bhutan, China, People's Republic of, India

Comment

Closely related to Nanorana liebigii (as Rana), according to the original publication. Original publication translated into English by Huang, Lathrop, and Murphy, 1998, Smithson. Herpetol. Inform. Serv., 118. Fei, 1999, Atlas Amph. China: 212-213, provided a brief account, map, and figure. Fei, Hu, Ye, and Huang, 2009, Fauna Sinica, Amph. 3: 1383-1387, provided an account (as Paa conaensis) for China, figures, and map, and included this species in their Paa liebigii group. Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 364, provided a brief account (as Paa conaensis) for China including photographs of specimens. Li, Zhao, and Dong, 2010, Amph. Rept. Tibet: 41-42, provided an account (as Paa conaensis) for Xizang, China. Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 442-443, provided an account (as Maculopaa conaensis), photographs, and a range map for China. Wangyal, 2013, J. Threatened Taxa, 5: 4777, provided records from Haa District, Bhutan. Saikia, Sinha, Dinesh, and Thakur, 2020, Rec. Zool. Surv. India, 120: 49–54, reported the species from Khremteng village, Tawang Township, Arunachal Pradesch, India. Che, Jiang, Yan, and Zhang, 2020, Amph. Rept. Tibet: 261–265, provided a detailed account for Tibet, China.

External links:

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For access to general information see Wikipedia
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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For additional information specific to China see Amphibia China
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.