Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Limnonectes tweediei (Smith, 1935)

Class: Amphibia > Order: Anura > Family: Dicroglossidae > Subfamily: Dicroglossinae > Genus: Limnonectes > Species: Limnonectes tweediei

Rana tweediei Smith, 1935, Bull. Raffles Mus., 10: 62. Holotype: BMNH 1934.5.21.1 (now reregistered as 1947.2.1.92) according to museum records,; given incorrectly as BMNH 1931.5.21.2.4 by Kiew, 1975 "1974", Malayan Nat. J., 28: 107. Type locality: "near the River Yum, Headwater Plus River, alt. 2,000 feet", Perak, Malaysia (Malaya).

Rana ulukalensis Nakatani, 1969, Acta Herpetol. Japon., 3: 10. Holotype: deposition not stated and presumed lost. Type locality: foot of Mount Ulu Kali, situated at the headwaters of Batang Kali, a river NNE of Kuala Lumpur, Malaysia. Synonymy by Das, 2010, Curr. Herpetol., Kyoto, 29: 35.

Rana ulikalensis Nakatani, 1969, Acta Herpetol. Japon., 3: 11. Incorrect original spelling according to Das, 2010, Curr. Herpetol., Kyoto, 29: 33.

Limnonectes (Limnonectes) tweediei — Dubois, 1987 "1986", Alytes, 5: 63.

English Names

Tweedie's Wart Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 100).

Distribution

Lowlands (up to 900 m elevation) of Peninsular Malaysia (states of Kedah, Perak, Selangor, Trengganu, and Pahang) and western Sumatra (Indonesia).

Geographic Occurrence

Natural Resident: Indonesia, Malaysia, Malaysia, West (Peninsular)

Comment

In the Limnonectes (Limnonectes) kuhlii group of Dubois, 1987 "1986", Alytes, 5: 63. Considered a synonym of Rana nitida by Kiew, 1975 "1974", Malayan Nat. J., 28: 107–109, but this synonymy was disputed by Dring, 1979, Bull. Brit. Mus. (Nat. Hist.), Zool., 34: 204–206, who posited that this species might be more closely related to Rana microdisca. See account by Bourret, 1942, Batr. Indochine: 284. See identification table by Manthey and Grossmann, 1997, Amph. Rept. Südostasiens: 84–86, to compare this species with other ranids of the Sunda Shelf region. See statement of geographic range, habitat, and conservation status in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 629. See comments on morphology and natural history in West Malaysia by Chan, Muin, Anuar, Andam, Razak, and Aziz, 2019, Check List, 15: 1055–1069.

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For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.