Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Bryophryne Hedges, Duellman, and Heinicke, 2008

Class: Amphibia > Order: Anura > Superfamily: Brachycephaloidea > Family: Strabomantidae > Subfamily: Holoadeninae > Genus: Bryophryne

11 species

Bryophryne Hedges, Duellman, and Heinicke, 2008, Zootaxa, 1737: 99. Type species: Phrynopus cophites Lynch, 1975, by original designation.

English Names

None noted.

Distribution

Amazonian slopes of the Cordillera Oriental in the Departamento Cusco, Peru, 2900–120 m elevation.

Comment

Partitioned from Phrynopus on phylogenetic grounds by Hedges, Duellman, and Heinicke, 2008, Zootaxa, 1737: 99, from which it is geographically separated by the Rio Apurimac valley according to Lehr and Catenazzi, 2009, S. Am. J. Herpetol., 4: 125–138, who mapped the type localities of the species know in August, 2009. Duellman and Lehr, 2009, Terrest.-breeding Frogs in Peru: 76–79, provided accounts. Pyron and Wiens, 2011, Mol. Phylogenet. Evol., 61: 543–583, suggested that Bryophryne is the sister taxon of Holoaden. This was rejected by González-Durán, Targino, Rada, and Grant, 2017, Zootaxa, 4243: 42–74.

Contained taxa (11 sp.):

Bryophryne abramalagae Lehr and Catenazzi, 2010

Bryophryne bakersfield Chaparro, Padial, Gutiérrez, and De la Riva, 2015

Bryophryne bustamantei (Chaparro, De la Riva, Padial, Ochoa, and Lehr, 2007)

Bryophryne cophites (Lynch, 1975)

Bryophryne hanssaueri Lehr and Catenazzi, 2009

Bryophryne nubilosus Lehr and Catenazzi, 2008

Bryophryne phuyuhampatu Catenazzi, Ttito, Diaz, and Shepack, 2017

Bryophryne quellokunka De la Riva, Chaparro, Castroviejo-Fisher, and Padial, 2017

Bryophryne tocra De la Riva, Chaparro, Castroviejo-Fisher, and Padial, 2017

Bryophryne wilakunka De la Riva, Chaparro, Castroviejo-Fisher, and Padial, 2017

Bryophryne zonalis Lehr and Catenazzi, 2009

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