Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Didynamipus Andersson, 1903

Class: Amphibia > Order: Anura > Family: Bufonidae > Genus: Didynamipus

1 species

Didynamipus Andersson, 1903, Verh. Zool. Bot. Ges. Wien, 53: 143. Type species: Didynamipus sjöstedti Andersson, 1903, by monotypy.

Atelophryne Boulenger, 1906 "1905", Ann. Mus. Civ. Stor. Nat. Genova, Ser. 3, 2: 159. Type species: Atelophryne minuta Boulenger, 1906, by monotypy. Synonymy by Mertens, 1965, Bonn. Zool. Beitr., 16: 19 (not Noble, 1924, Bull. Am. Mus. Nat. Hist., 49: 147-347, as reported by Grandison, 1981, Monit. Zool. Ital., N.S., Suppl., 15: 188).

English Names

Four-digit Toads (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 44).

Dwarf Toads (Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 94).

Distribution

Bioko (Equatorial Guinea) and extreme southwestern Cameroon to Nigeria.

Comment

The phylogenetic relationships of this genus were discussed by Grandison, 1981, Monit. Zool. Ital., N.S., Suppl., 15: 187-215. Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 129, suggested on the basis of DNA sequence evidence that Didynamipus is most closely related to Nectophrynoides among the exemplars studied by those authors. Smith and Chiszar, 2006, Herpetol. Conserv. Biol., 1: 6-8, implied that this taxon should be considered a subgenus of Bufo; see comment under Bufonidae. Van Bocxlaer, Biju, Loader, and Bossuyt, 2009, BMC Evol. Biol., 9 (e131): 1-10, did not address this genus. Van Bocxlaer, Loader, Roelants, Biju, Menegon, and Bossuyt, 2010, Science, 327: 679-682, suggested that Didynamipus phylogenetically isolated, in the neighborhood of Schismaderma, Churamiti, Nectophrynoides, but not demonstratably close to them. Channing, Rödel, and Channing, 2012, Tadpoles of Africa: 142, suggested that the species is a direct-developer.

Contained taxa (1 sp.):

Didynamipus sjostedti Andersson, 1903

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