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Heinicke, M. P., W. E. Duellman, L. Trueb, D. B. Means, R. D. MacCulloch, and S. B. Hedges. 2009. A new frog family (Anura: Terrarana) from South America and an expanded direct-developing clade revealed by molecular phylogeny. Zootaxa 2211: 1–35.