Bibliography

Guayasamin, J. M., E. Bonaccorso, W. E. Duellman, and L. A. Coloma. 2010. Genetic differentiation in the nearly extinct harlequin frogs (Bufonidae: Atelopus), with emphasis on the Andean Atelopus ignescens and A. bomolochos species complexes. Zootaxa 2574: 55–68.