Tylototriton hainanensis Fei, Ye, and Yang, 1984

Class: Amphibia > Order: Caudata > Family: Salamandridae > Subfamily: Pleurodelinae > Genus: Tylototriton > Species: Tylototriton hainanensis

Tylototriton hainanensis Fei, Ye, and Yang, 1984, Acta Zool. Sinica, 30: 85-89. Holotype: CIB 64III1379, by original designation. Type locality: "Nalong, Wuzhi Shan, alt. 770 m", Hainan, China.

Pleurodeles (Tylototrion) hainanensisRisch, 1985, J. Bengal Nat. Hist. Soc., N.S.,, 4: 142.

Pleurodeles (Echinotriton) hainanensisDubois, 1987 "1986", Alytes, 5: 11.

Tylototriton (Yaotriton) hainanensisDubois and Raffaëlli, 2009, Alytes, 26: 68.

Yaotriton hainanensis —  Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 91; Dubois, Ohler, and Pyron, 2021, Megataxa, 5: 408. 

English Names

Hainan Knobby Newt (Fei, 1999, Atlas Amph. China: 40).

Hainan Crocodile Newt (Raffaëlli, 2022, Salamanders & Newts of the World: 287).

Distribution

Biasha, Leong, Lingshui and Qiongzhong, Hainan Island, China, 770–970 m elevation. See comment. 

Geographic Occurrence

Natural Resident: China, People's Republic of

Endemic: China, People's Republic of

Comment

See accounts by Ye, Fei, and Hu, 1993, Rare and Economic Amph. China 74, and Fei, 1999, Atlas Amph. China: 40. Synonymy with Tylototrion asperrimus by Zhao, Hu, Jiang, and Yang, 1988, Studies on Chinese Salamanders: 63; this rejected by Ye, Fei, and Hu, 1993, Rare and Economic Amph. China 74. In the Tylototriton asperrimus group of Fei, Ye, Huang, Jiang, and Xie, 2005, in Fei et al. (eds.), Illust. Key Chinese Amph.: 42 (although they only addressed Chinese species). See comments under Tylototriton and Tylototriton asperrimus. Fei, Hu, Ye, and Huang, 2006, Fauna Sinica, Amph. 1: 261–265, provided an account. See photograph, map, description of geographic range and habitat, and conservation status in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 607. Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 76, provided a brief account including photographs. Shi, 2011, Amph. Rept. Fauna Hainan: 36–38, provided an account. Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 91, provided an account, photographs, and a map. Raffaëlli, 2013, Urodeles du Monde, 2nd ed.: 186, provided a brief account, photograph, and map. Sparreboom, 2014, Salamanders Old World: 361–362, reviewed the biology, characteristics, distribution, reproduction, and conservation of the species. Fei and Ye, 2016, Amph. China, 1: 301–303, provided an account (as Yaotriton hainanensis), photographs, and range map.  Bernardes, Le, Nguyen, Pham, Pham, Nguyen, Rödder, Bonkowski, and Ziegler, 2020, ZooKeys, 935: 121–164, provided a range map and mapped a population in western Guizhou, China, as Tylototriton cf. hainanensis, implying an unnamed species. In the Tylototriton (Yaotritonasperrimus species group of Poyarkov, Nguyen, and Arkhipov, 2021, Taprobanica, 10: 4–22, who discussed phylogenetics. In the Tylototriton asperrimus group of Lyu, Wang, Zeng, Zhou, Qi, Wan, Li, and Wang, 2021, Vert. Zool., Senckenberg, 71: 697–710, who discussed phylogenetics. Raffaëlli, 2022, Salamanders & Newts of the World: 287–288, provided an account, summarizing systematics, life history, population status, and distribution (including a polygon map). 

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