Tylototriton asperrimus Unterstein, 1930

Class: Amphibia > Order: Caudata > Family: Salamandridae > Subfamily: Pleurodelinae > Genus: Tylototriton > Species: Tylototriton asperrimus

Tylototriton asperrimus Unterstein, 1930, Sitzungsber. Ges. Naturforsch. Freunde Berlin, 1930: 314. Syntypes: ZMB (2 specimens); ZMB 34089 considered holotype (lectotype designation by implication) by Bauer, Good, and Günther, 1993, Mitt. Zool. Mus. Berlin, 69: 298. Type locality: "Kwangsi"; corrected to "Yao Shan [= Mt. Yao], Kwangsi Prov. [= Guangxi Zhuang Autonomous Region], China" by Bauer, Good, and Günther, 1993, Mitt. Zool. Mus. Berlin, 69: 298. Fan, 1931, Bull. Dept. Biol. Coll. Sci. Sun Yatsen Univ., 11: 1–154, provided evidence that the type locality is or near to the town of Loshiang.

Tylototriton (Echinotriton) asperrimusZhao and Hu, 1984, Stud. Chinese Tailed Amph.: 19.

Tylototriton asperrimus asperrimusFei, Ye, and Yang, 1984, Acta Zool. Sinica, 30: 89.

Pleurodeles (Tylototrion) asperrimus asperrimusRisch, 1985, J. Bengal Nat. Hist. Soc., N.S.,, 4: 142.

Pleurodeles (Echinotriton) asperrimusDubois, 1987 "1986", Alytes, 5: 11.

Echinotriton asperrimusZhao, 1990, in Zhao (ed.), From Water onto Land: 219; Nguyen, Ho, and Nguyen, 2005, Checklist Amph. Rept. Vietnam: 9.

Echinotriton asperrimus asperrimusZhao and Adler, 1993, Herpetol. China: 112.

Tylototriton asperrimusNussbaum, Brodie, and Yang, 1995, Herpetologica, 51: 265.

Tylototriton (Yaotriton) asperrimusDubois and Raffaëlli, 2009, Alytes, 26: 68.

Yaotriton asperrimus — Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 88.

English Names

Mount Yao Newt (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 35).

Black Knobby Newt (Fei, 1999, Atlas Amph. China: 40).

Granular Newt (Nguyen, Ho, and Nguyen, 2005, Checklist Amph. Rept. Vietnam: 9).


Guangxi (Darongshan, Huanjiang, Longsheng, Xincheng, Yaoshan, Yulin, and Napo counties), Guangdong (Xinyi County), and Guizhou (Libo), China, 1320 to 1400 m elevation; Mau Son and Lang Son areas of Ha Giang and Lao Cai provinces in northern Vietnam, 1400–1700 m elevation.


See accounts by Bourret, 1942, Batr. Indochine: 146-148; Fang and Chang, 1932, Sinensia, Nanking, 2: 117-121; Ye, Fei, and Hu, 1993, Rare and Economic Amph. China 72; and Fei, 1999, Atlas Amph. China: 40. See also Liu, Hu, Fei, and Huang, 1973, Acta Zool. Sinica, 19: 394. Deng and Yu, 1984, Acta Herpetol. Sinica, Chengdu, N.S.,, 3 (2): 75-77, recognize a subspecies in Sichuan. Nussbaum, Brodie, and Yang, 1995, Herpetologica, 51: 265, discussed reasons for transferring this species from Echinotriton to TylototritonZhang and Wen, 2000, Amph. Guangxi: 21, provided an account for population in Guangxi, China.Bain and Nguyen, 2004, Am. Mus. Novit., 3453: 9, provided the record for Ha Giang Province, Vietnam (although these records may now be reassigned to another species). Nguyen, Ho, and Nguyen, 2005, Checklist Amph. Rept. Vietnam: 9, provided specific records for Vietnam and (p. 147) provided a photograph. In the Tylototriton asperrimus group of Fei, Ye, Huang, Jiang, and Xie, 2005, in Fei et al. (eds.), Illust. Key Chinese Amph.: 42 (although they only addressed Chinese species). Böhme, Schöttler, Nguyen, and Köhler, 2005, Salamandra, 41:218-219, discussed previous confusion in Vietnam of Tylototriton asperrimus and Tylototriton vietnamensis and noted that the record for Tam Dao, northern Vietnam, may actually be referrable to Tylototriton hainanensis. Fei, Hu, Ye, and Huang, 2006, Fauna Sinica, Amph. 1: 257-261, provided an account. See statement of geographic range, habitat, and conservation status in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 643. Nguyen, Nguyen, Hồ, Lê, and Nguyen, 2009, Tạp chí Công nghệ Sinh học, 7: 325-333, discussed the species in Vietnam. Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 72-73, provided a brief account including photographs of specimens and habitat. Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 88–89, provided an account, photographs, and a map that excluded Vietnam from the range. A record for Thuong Tien Nature Reserve, Hoa Binh Province, in northern Vietnam, reported by Luu, Le, Do, Hoang, Nguyen, Bonkowski, and Ziegler, 2014, Herpetol. Notes, 7: 51–58. Raffaëlli, 2013, Urodeles du Monde, 2nd ed.: 186–186, provided a brief account, photograph, and map. Fei and Ye, 2016, Amph. China, 1: 292–294, provided an account (as Yaotriton apserrimus), photographs, and range map. 

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