Calotriton asper (Dugès, 1852)

Class: Amphibia > Order: Caudata > Family: Salamandridae > Subfamily: Pleurodelinae > Genus: Calotriton > Species: Calotriton asper

Triton glacialis Philippe, 1847, Séances Acad. Sci. Lettr. Montpellier, 1847: 20. Type(s): Not stated or known to exist. Type locality: Lac Bleu, Pyrenees [France]. Nomen nudum. Synonymy by Mertens and Müller, 1928, Abh. Senckenb. Naturforsch. Ges., 41: 10.

Hemitriton asper Dugès, 1852, Ann. Sci. Nat., Paris, Ser. 3, 17: 266. Syntypes: MNHNP 1082 (2 specimens), according to Guibé, 1950 "1948", Cat. Types Amph. Mus. Natl. Hist. Nat.: 7. These numbered MNHNP 4738 and 4738A according to Thireau, 1986, Cat. Types Urodeles Mus. Natl. Hist. Nat., Rev. Crit.: 15. Also including MDUG HE 1219-21, according to Flores-Villela, Ríos-Muñoz, Magaña-Cota, and Quezadas-Tapia, 2016, Zootaxa, 4092: 39. Type locality: "Eaux-Bonnes", Pyrenees, France.

Triton repandus Dugès, 1852, Ann. Sci. Nat., Paris, Ser. 3, 17: 256. Substitute name for Hemitriton asper attributed to Duméril and Valenciennes.

Hemitriton cinereus Dugès, 1852, Ann. Sci. Nat., Paris, Ser. 3, 17: 263. Syntypes: MNHNP 4741 (2 specimens), according to Guibé, 1950 "1948", Cat. Types Amph. Mus. Natl. Hist. Nat.: 7, and Thireau, 1986, Cat. Types Urodeles Mus. Natl. Hist. Nat., Rev. Crit.: 26. Type locality: Pyrenees. Synonymy by Boulenger, 1882, Cat. Batr. Grad. Batr. Apoda Coll. Brit. Mus., Ed. 2: 24.

Hemitriton rugosus Dugès, 1852, Ann. Sci. Nat., Paris, Ser. 3, 17: 264. Holotype: MNHNP 4740, according to Guibé, 1950 "1948", Cat. Types Amph. Mus. Natl. Hist. Nat.: 8. Type locality: Pyrenees? Synonymy by Boulenger, 1882, Cat. Batr. Grad. Batr. Apoda Coll. Brit. Mus., Ed. 2: 24.

Hemitriton punctulatus Dugès, 1852, Ann. Sci. Nat., Paris, Ser. 3, 17: 265. Syntypes: MNHNP 4739 (2 specimens), according to Guibé, 1950 "1948", Cat. Types Amph. Mus. Natl. Hist. Nat.: 8, and Thireau, 1986, Cat. Types Urodeles Mus. Natl. Hist. Nat., Rev. Crit.: 60. Type locality: "Eaux-Bonnes (Pyrénées)", France. Synonymy by Boulenger, 1882, Cat. Batr. Grad. Batr. Apoda Coll. Brit. Mus., Ed. 2: 24.

Hemitriton bibronii Dugès, 1852, Ann. Sci. Nat., Paris, Ser. 3, 17: 266. Syntypes: MNHNP 1082 (4 specimens), according to Guibé, 1950 "1948", Cat. Types Amph. Mus. Natl. Hist. Nat.: 7; MNHNP 1082, 1082A-C according to Thireau, 1986, Cat. Types Urodeles Mus. Natl. Hist. Nat., Rev. Crit.: 18. Type locality: Pyrenees. Synonymy by Boulenger, 1882, Cat. Batr. Grad. Batr. Apoda Coll. Brit. Mus., Ed. 2: 24.

Triton pyrenaeus Duméril, Bibron, and Duméril, 1854, Erp. Gen., 9: 139. Holotype: MNHNP 5337, according to Guibé, 1950 "1948", Cat. Types Amph. Mus. Natl. Hist. Nat.: 8, Thireau, 1986, Cat. Types Urodeles Mus. Natl. Hist. Nat., Rev. Crit.: 61. Type locality: Pyrenees, France. Synonymy by Boulenger, 1882, Cat. Batr. Grad. Batr. Apoda Coll. Brit. Mus., Ed. 2: 24.

Triton rugosusDuméril, Bibron, and Duméril, 1854, Erp. Gen., 9: 150.

Triton cinereusDuméril, Bibron, and Duméril, 1854, Erp. Gen., 9: 151.

Triton repandusDuméril, Bibron, and Duméril, 1854, Erp. Gen., 9: 151.

Triton puncticulatusDuméril, Bibron, and Duméril, 1854, Erp. Gen., 9: 140. Incorrect subsequent spelling of punctulatus.

Triton bibroniDuméril, Bibron, and Duméril, 1854, Erp. Gen., 9: 153.

Calotriton punctulatusGray, 1858, Proc. Zool. Soc. London, 1858: 139.

Euproctus pyrenaeusLataste, 1878, Rev. Internatl. Sci., 1st year, Paris, 2: 495.

Molge asperaBoulenger, 1882, Cat. Batr. Grad. Batr. Apoda Coll. Brit. Mus., Ed. 2: 24.

Euproctus asperaCamerano, 1885, Mem. Accad. Sci. Torino, Ser. 2, 36: 419.

Euproctus asper var. rugosa Bedriaga, 1895, Proc. Zool. Soc. London, 1895: 160. Syntypes: MNHNP and ZMB, according to XXX. Type locality: Region of the Pyrennees. Synonymy by Wolterstorff, 1925, Abh. Ber. Mus. Nat. Heimatkd. Magdeburg, 4: 64.

Triton asperWolterstorff, 1902, Triton der Untergattung Euproctus: 12. Schreiber, 1912, Herpetol. Eur., Ed. 2: 48.

Molge asperaMaluquer, 1916, Butll. Inst. Catalana Hist. Nat., Ser. 2, 13: 58.

Diemictylus asperFowler and Dunn, 1917, Proc. Acad. Nat. Sci. Philadelphia, 69: 28.

Molge bolivari Boscá, 1918, Bol. R. Soc. Esp. Hist. Nat., 18: 58. Type(s): Not designated although presumably in MNCN. Type locality: "valle deTena, en la vertiente meridional de los Pirineos aragoneses a 1.600 ó más metros de altur sobre el nivel del more, donde se halla el sanatorio de Panticosa (Huesca)", Spain. Synonymy by Mertens and Wermuth, 1960, Amph. Rept. Europas: 17.

Euproctus asperWolterstorff, 1925, Abh. Ber. Mus. Nat. Heimatkd. Magdeburg, 4: 61.

Euproctus asper forma d'Onceti Wolterstorff, 1925, Abh. Ber. Mus. Nat. Heimatkd. Magdeburg, 4: 66. Holotype: MM 97/24, presumably destroyed in W.W.II. Type locality: "Lac d'Oncet bei Bangnères de Bigorre", Pyrenees, France.

Euproctus asper forma castelmouliensis Wolterstorff, 1925, Abh. Ber. Mus. Nat. Heimatkd. Magdeburg, 4: 66. Syntypes: MM 99/24 (2 specimens) and 100/24 (1 specimen), destroyed in WWII according to Brame, 1972, Checklist Living & Fossil Salamand. World (Unpubl. MS): 49; SMF 1167 regarded as lectotype (presumably exchanged from MM) by Mertens, 1967, Senckenb. Biol., 48(A): 37. See comments by Glaw and Franzen, 2006, Spixiana, München, 29: 157. Type locality: "Bergbach und seine Umbegung, liegt ebenfalls bei Bagnères de Bigorre in ca. 7-800 m Meereshöhe", Pyrenees, France.

Euproctus asper forma peyreladensis Wolterstorff, 1925, Abh. Ber. Mus. Nat. Heimatkd. Magdeburg, 4: 71. Holotype: MM 110/24 (Destroyed in W.W.II), according to XXX. Type locality: "Bagnères de Bigorre aus dem Lac de Peyrelade", Pyrenees, France.

Euproctus (Hemitriton) asperWolterstorff, 1925, Abh. Ber. Mus. Nat. Heimatkd. Magdeburg, 4: 296.

Euproctus asper asperMertens and Müller, 1928, Abh. Senckenb. Naturforsch. Ges., 41: 9.

Euproctus asperWolterstorff and Herre, 1935, Arch. Naturgesch., Leipzig, N. F., 4: 224.

Euproctus asper castelmouliensisMertens and Müller, 1940, Abh. Senckenb. Naturforsch. Ges., 451: 8.

Calotriton asperCarranza and Amat, 2005, Zool. J. Linn. Soc., 145: 555.

English Names

Pyrenean Newt (Bedriaga, 1895, Proc. Zool. Soc. London, 1895: 160; Martínez Rica, 1997, in Gasc et al. (eds.), Atlas Amph. Rept. Eur.: 54).

Pyrenean Brook Salamander (Arnold and Burton, 1978, Field Guide Rept. Amph. Brit. Eur.: 38; Ananjeva, Borkin, Darevsky, and Orlov, 1988, Dict. Amph. Rept. Five Languages: 32; Stumpel-Rienks, 1992, Ergänzungsband Handbuch Rept. Amph. Eur., Trivialnamen der Herpetofauna Eur.: 48).

Pyrenean Mountain Salamander (Steward, 1969, Tailed Amph. Eur.: 39).

Pyrenean Mountain Newt (Ananjeva, Borkin, Darevsky, and Orlov, 1988, Dict. Amph. Rept. Five Languages: 32; Stumpel-Rienks, 1992, Ergänzungsband Handbuch Rept. Amph. Eur., Trivialnamen der Herpetofauna Eur.: 48).

Pyrenees Mountain Salamander (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 35).

Pyrenees Mountain Newt (Thorn and Raffaëlli, 2000, Salamand. Ancien Monde: 350).

Pyreneen Salamander (Hellmich, 1962, Rept. Amph. Eur.: 52).

Distribution

Pyrenees Mountains of France, Spain, and Andorra, 175-3000 m elevation.

Comment

Thorn and Raffaëlli, 2000, Salamand. Ancien Monde: 350-357, provided an account. See also discussion of relevant literature and distribution by Martínez Rica, 1997, in Gasc et al. (eds.), Atlas Amph. Rept. Eur.: 54-55. Clergue-Gazeau, 1999, in Grossenbacher and Thiesmeier (eds.), Handbuch Rept. Amph. Eur., 4(1): 251-269, provided a review of the biology. Barbadillo-Escrivá, 1987, Guia INCAFO Anf. Rept. Peninsula Iberica: 100-105, provided an account for the Iberian populations. Obst IN Engelmann, Fritzsche, Günther, and Obst, 1993, Lurche Kriechtiere Eur.: 68-70, provided a brief account, figure, and map. Llorente, Montori, Santos, and Carretero, 1995, Atlas Amf. Rept. Catalunya Andorra: 46-48, provided an account and detailed range map for Catalunya. Raffaëlli, 2007, Les Urodèles du Monde: 121, provided a brief account, map, and photograph. Poschadel and Plath, 2004, Elaphe, 12: 59-67, discussed habitat preference and interpopulation relationships. Montori, Llorente, and García-París, 2008, Zootaxa, 1945: 39-50, reported on geographic variation in allozymes. See statement of geographic range, habitat, and conservation status in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 641-642. Nöllert and Nöllert, 1992, Die Amph. Eur.: 159-162, provided a brief account (as Euproctus asper) and polygon map. Rivera, Escoriza, Maluquer-Margalef, Arribas, and Carranza, 2011, Amf. Rept. Catalunya: 43-45, provided a brief account for northeastern Spain, Andorra, and adjacent France.

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