Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Hylarana spinulosa (Smith, 1923)

Class: Amphibia > Order: Anura > Family: Ranidae > Genus: Hylarana > Species: Hylarana spinulosa

Rana (Hylarana) spinulosa Smith, 1923, J. Nat. Hist. Soc. Siam, 6: 207. Holotype: BMNH (formerly M. Smith 6889). Type locality: "Tun-fao, Kachek river [= Jiaji He], altitude 200 metres", Hainan I., China.

Hylarana spinulosa — Bourret, 1942, Batr. Indochine: 98; Song, Jang, Zou, and Shi, 2002, Herpetol. Sinica, 9: 71; Chen, Murphy, Lathrop, Ngo, Orlov, Ho, and Somorjai, 2005, Herpetol. J., 15: 237, by implication; Che, Pang, Zhao, Wu, Zhao, and Zhang, 2007, Mol. Phylogenet. Evol., 43: 3, by implication; Dubois, Ohler, and Pyron, 2021, Megataxa, 5: 423.

Hylarana (Hylarana) spinulosa — Fei, Ye, and Huang, 1990, Key to Chinese Amph.: 140; Fei, 1999, Atlas Amph. China: 176.

Rana (Sylvirana) spinulosa — Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 326.

Sylvirana spinulosa — Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 370; Fei, Ye, and Jiang, 2010, Herpetol. Sinica, 12: 34; Oliver, Prendini, Kraus, and Raxworthy, 2015, Mol. Phylogenet. Evol., 90: 191.

Hylarana (Sylvirana) spinulosa — Fei, Hu, Ye, and Huang, 2009, Fauna Sinica, Amph. 3: 1143.

English Names

Spiny Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 109).

Fine-spined Frog (Fei, 1999, Atlas Amph. China: 176).

Distribution

Southern and southwestern Hainan Island, China, 80–840 m elevation. See comment.

Geographic Occurrence

Natural Resident: China, People's Republic of

Endemic: China, People's Republic of

Comment

Prior to the revision of Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297, placed in the Section Hylarana, subsection Hydrophylax, subgenus Sylvirana of Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 326. See Bourret, 1942, Batr. Indochine: 320–321; Liu and Hu, 1961, Tailless Amph. China: 190–191; and Fei, 1999, Atlas Amph. China: 176–177. In the Hylarana (Hylarana) latouchii group of Fei, Ye, Huang, Jiang, and Xie, 2005, in Fei et al. (eds.), Illust. Key Chinese Amph.: 117. Yang and Zhao, 2005, Sichuan J. Zool., 24: 309–311, provided a specific location and habitat. See photograph, map, description of geographic range and habitat, and conservation status (as Rana spinulosa) in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 512. Fei, Hu, Ye, and Huang, 2009, Fauna Sinica, Amph. 3: 1143–1148, provided an account, figures, and map for China and placed it in their Hylarana (Sylvirana) latouchii group. Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 319, provided a brief account including photographs. Shi, 2011, Amph. Rept. Fauna Hainan: 116–119, provided an account as Rana spinulosa. Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 359, provided an account (as Sylvirana spinulosa), photographs, and a range map. Gawor, Pham, Nguyen, Nguyen, Schmitz, and Ziegler, 2016, Salamandra, 52: 28–29, reported on an apparently unnamed "Hylarana sp." from Quang Ninh Province, in northeastern Vietnam that resembled this species.

External links:

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For access to general information see Wikipedia
For additional sources of general information from other websites search Google
For access to relevant technical literature search Google Scholar
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For additional information specific to China see Amphibia China
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.