Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Hylarana amnicola Perret, 1977

Class: Amphibia > Order: Anura > Family: Ranidae > Genus: Hylarana > Species: Hylarana amnicola

Hylarana amnicola Perret, 1977, Rev. Suisse Zool., 84: 855. Holotype: MHNG 1551.43, by original designation. Type locality: "Ilanga, Eséka, Cameroun méridional".

Rana amnicola — Perret, 1985, in Frost (ed.), Amph. Species World: 480.

Rana (Hylarana) amnicola — Dubois, 1987 "1986", Alytes, 5: 42, by implication.

Rana (Amnirana) amnicola — Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 324.

Amnirana amnicola — Channing, 2001, Amph. Cent. S. Afr.: 266, by implication; Fei, Ye, and Jiang, 2010, Herpetol. Sinica, 12: 33. See comment under Ranidae record.

Hydrophylax amnicola — Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 368.

Hylarana amnicola — Che, Pang, Zhao, Wu, Zhao, and Zhang, 2007, Mol. Phylogenet. Evol., 43: 1-13, by implication; Dubois, Ohler, and Pyron, 2021, Megataxa, 5: 461, by implication.

Amnirana amnicola — Oliver, Prendini, Kraus, and Raxworthy, 2015, Mol. Phylogenet. Evol., 90: 187.

English Names

Ilanga Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 106).

Stream White-lipped Frog (Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 390).

Distribution

Central and western Cameroon, Equatorial Guinea, Gabon, and northwestern Rep. Congo; reported from northeastern Dem. Rep. Congo (see comment).

Geographic Occurrence

Natural Resident: Cameroon, Congo, Democratic Republic of the, Congo, Republic of the, Equatorial Guinea, Gabon

Comment

In the Rana (Amnirana) albolabris group of Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 324. De la Riva, 1994, Rev. Esp. Herpetol., 8: 135, provided a record for Equatorial Guinea. Frétey and Blanc, 2002 "2001", Bull. Soc. Zool. France, 126: 385, reported this species from Gabon. Lasso, Rial, Castroviejo, and De la Riva, 2002, Graellsia, 58: 21-34, provided notes (as Amnirana amnicola) on ecological distribution in Equatorial Guinea. Penner and Rödel, 2007, in Butynski and McCullough (eds.), RAP Bull. Biol. Assessment, 46: 37-41, provided a record for north-central Dem. Rep. Congo, although this record is out of range according to the map provided by Jongsma, Barej, Barratt, Burger, Conradie, Ernst, Greenbaum, Hirschfeld, Leaché, Penner, Portik, Zassi-Boulou, Rödel, and Blackburn, 2018, Mol. Phylogenet. Evol., 120: 275. Jongsma, Tobi, Dixon-MacCallum, Bamba-Kaya, Yoga, Mbega, Mve Beh, Emrich, and Blackburn, 2017, Amph. Rept. Conserv., 11 (1: e144): 16, provided records for southeastern Gabon and briefly discussed habitat and range. Deichmann, Mulcahy, Vanthomme, Tobi, Wynn, Zimkus, and McDiarmid, 2017, PLoS One, 12 (11: e0187283): 24, reported molecular evidence in support of an unnamed lineage (their Amnirana cf. amnicola") in Rep. Congo. Dewynter, Frétey, Jongsma, Bamba-Kaya, and Pauwels, 2018, Cah. Fondation Biotope, 18: 16, provided a brief account and photographs from the Monts Birougou area of Gabon. Mali, Banda, Chifundera, Badjedjea, Sebe, Lokasola, Ewango, Tungaluna, and Akaibe, 2019, Am. J. Zool., 2: 38–43, reported the species from the Okapi Wildlife Reserve, northeastern Dem. Rep. Congo, although this identification requires molecular confirmation. Dewynter and Frétey, 2019, Cah. Fondation Biotope, 27: 40, summarized the literature for Gabon and provided photographs (p. 66). Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 392–393, provided a brief account, photograph, and range map. See brief account, photographs, and range map for Equatorial Guinea in Sánchez-Vialas, Calvo-Revuelta, Castroviejo-Fisher, and De la Riva, 2020, Proc. California Acad. Sci., Ser. 4, 66: 137–230.

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For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.