Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Leptodactylus rugosus Noble, 1923

Class: Amphibia > Order: Anura > Family: Leptodactylidae > Subfamily: Leptodactylinae > Genus: Leptodactylus > Species: Leptodactylus rugosus

Leptodactylus rugosus Noble, 1923, Zoologica, New York, 3: 297. Holotype: AMNH 1169, by original designation. Type locality: "near Kaieteur Falls, British Guiana [=Guyana]".

English Names

Guyana White-lipped Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 82).

Rugose Thin-toed Frog (Kok and Kalamandeen, 2008, Intr. Taxon. Amph. Kaieteur Natl. Park: 228).

Distribution

Southern Venezuela (Bolívar) east to western Guyana; likely in intervening adjacent Brazil (see comment).

Geographic Occurrence

Natural Resident: Brazil, Guyana, Venezuela

Comment

In the Leptodactylus pentadactylus group of Heyer, 1972, Contrib. Sci. Nat. Hist. Mus. Los Angeles Co., 231: 1-8, and Heyer, 1979, Smithson. Contrib. Zool., 301: 1-43. See account in Donnelly and Myers, 1991, Am. Mus. Novit., 3017D: 21-23. See also Heyer, 1995, Proc. Biol. Soc. Washington, 108: 695-716. Duellman, 1997, Sci. Pap. Nat. Hist. Mus. Univ. Kansas, 2: 25, commented on the southeastern Venezuela population and described the larva and natural history. See Heyer and Thompson, 2000, Cat. Am. Amph. Rept., 708: 1-5, for review. Myers and Donnelly, 2008, Bull. Am. Mus. Nat. Hist., 308: 82-88, reported on material from Auyantepui, Venezuela. Kok and Kalamandeen, 2008, Intr. Taxon. Amph. Kaieteur Natl. Park: 228-229, provided an account. MacCulloch and Lathrop, 2009, R. Ontario Mus. Contrib. Sci., 4: 15, commented on specimens from Mount Ayanganna, Guyana. See Cole, Townsend, Reynolds, MacCulloch, and Lathrop, 2013, Proc. Biol. Soc. Washington, 125: 427, for brief account and records for Guyana. Señaris, Lampo, Rojas-Runjaic, and Barrio-Amorós, 2014, Guía Ilust. Anf. Parque Nac. Canaima: 216–217, provided a photograph and a brief account for the Parque Nacional de Canaima, Venezuela. In the Leptodactylus pentadactylus species group of de Sá, Grant, Camargo, Heyer, Ponssa, and Stanley, 2014, S. Am. J. Herpetol., 9(Spec. Issue 1): 1–123, and who provided a summary of relevant literature (adult and larval morphology, identification, advertisement call, and range) on pp. 60–61. See Barrio-Amorós, Rojas-Runjaic, and Señaris, 2019, Amph. Rept. Conserv., 13 (1: e180): 97, for comments on range and literature. Segalla, Berneck, Canedo, Caramaschi, Cruz, Garcia, Grant, Haddad, Lourenço, Mângia, Mott, Nascimento, Toledo, Werneck, and Langone, 2021, Herpetol. Brasil., 10: 190, included this species in the Brazilian fauna, but DRF is unaware of any published localities.

External links:

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For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.