Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Pleurodema cinereum Cope, 1878

Class: Amphibia > Order: Anura > Family: Leptodactylidae > Subfamily: Leiuperinae > Genus: Pleurodema > Species: Pleurodema cinereum

Pleurodema cinereum Cope, 1878 "1877", Proc. Am. Philos. Soc., 17: 40. Holotype: "No. 137", presumably deposited in ANSP or Philadelphia Museum, but location currently unknown. Type locality: "Juliaca at 12,500 feet", Peru.

Paludicola cinerea — Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 232.

Paludicola alpina Andersson, 1906, Ark. Zool., 3(12): 6. Syntypes: 2 specimens (Stockholm Högskola collection, now ZIUS?). Type locality: "Casabindo, Puna de Jujuy, Andes of Argentina, 3500 m". Synonymy by Parker, 1927, Ann. Mag. Nat. Hist., Ser. 9, 20: 470.

Pleurodema alpina — Nieden, 1923, Das Tierreich, 46: 497.

Pleurodema cinerea — Nieden, 1923, Das Tierreich, 46: 498; Parker, 1927, Ann. Mag. Nat. Hist., Ser. 9, 20: 470.

Paludicola (Pleurodema) alpina — Andersson, 1939 "1938", Ark. Zool., 30(23): 11.

Pleurodema cinerea cinerea — Cei, 1956, Invest. Zool. Chilen., 3: 51.

Pleurodema cinerea alpina — Cei, 1956, Invest. Zool. Chilen., 3: 52.

Pleurodema alpina — Gallardo, 1968, Physis, Buenos Aires, 28: 142.

Pleurodema cinereum — Duellman, 1993, Univ. Kansas Mus. Nat. Hist. Spec. Publ., 21: 128; De la Riva and Gonzales-Álvarez, 1998, Alytes, 16: 68. Justified emendation.

English Names

Juliaca Four-eyed Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 84).

Distribution

Northwestern Argentina through Bolivia to southeastern Peru along the Andes, 1000–4200 m elevation.

Geographic Occurrence

Natural Resident: Argentina, Bolivia, Peru

Comment

See comment under Pleurodema borellii. See account by Cei, 1980, Monit. Zool. Ital., N.S., Monogr., 2: 361–366. Márquez, De la Riva, and Bosch, 1995, J. Zool., London, 237: 313–336, reported on vocalization in Bolivia. Köhler, 2000, Bonn. Zool. Monogr., 48: 139, provided a brief account. Ferraro and Casagranda, 2009, Zootaxa, 2024: 33–55, provided a detailed discussion of range in Argentina. Ferraro, Blotto, Baldo, Barrasso, Barrionuevo, Basso, Cardozo, Cotichelli, Faivovich, Pereyra, and Lavilla, 2018, in Vaira, Akmentins, and Lavilla (eds.), Cuad. Herpetol., 32 (Supl. 1): 17–19, noted that the taxonomic status of this species with respect to Pleurodema borellii remains problematic.

External links:

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For access to general information see Wikipedia
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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.