Hyperolius viridiflavus (Duméril and Bibron, 1841)

Class: Amphibia > Order: Anura > Family: Hyperoliidae > Genus: Hyperolius > Species: Hyperolius viridiflavus

Eucnemis viridi-flavus Duméril and Bibron, 1841, Erp. Gen., 8: 528. Syntypes: MNHNP 412 and 4599 (according to Guibé, 1950 "1948", Cat. Types Amph. Mus. Natl. Hist. Nat.: 57) and NHMW 22896 (according to Häupl and Tiedemann, 1978, Kat. Wiss. Samml. Naturhist. Mus. Wien, 2: 16, and Häupl, Tiedemann, and Grillitsch, 1994, Kat. Wiss. Samml. Naturhist. Mus. Wien, 9: 21). The status of these is questionable as measurements are provided for only one specimen in the original publication (holotype?) and Laurent, 1951, Ann. Soc. R. Zool. Belg., 82: 40, reported only MNHNP 412 (2 specimens) as syntypes. Type locality: "Abyssinie".

Hyperolius viridiflavusGünther, 1859 "1858", Cat. Batr. Sal. Coll. Brit. Mus.: 85; Noble, 1924, Bull. Am. Mus. Nat. Hist., 49: 252, 330; Ahl, 1931, Das Tierreich, 55: 319.

Rappia bayoni Boulenger, 1911, Ann. Mus. Civ. Stor. Nat. Genova, Ser. 3, 5: 168. Syntypes: MSNG and including MCZ 3232, according to Barbour and Loveridge, 1929, Bull. Mus. Comp. Zool., 69: 333; MSNG 29404A designated lectotype by Capocaccia, 1957, Ann. Mus. Civ. Stor. Nat. Genova, Ser. 3, 69: 218. Type localities: "Entebbe", "Bussu", "Bululo", "Mbale", "Jinja", and "Kabulamuliro", Uganda. Synonymy (with Hyperolius viridiflavus) by Laurent, 1950, Explor. Parc Natl. Albert, Miss. G.F. de Witte (1933–1935), 64: 76. Considered a subspecies of Hyperolius viridiflavus by Laurent, 1961, Rev. Zool. Bot. Afr., 64: 91. Synonymy by Schiøtz, 1971, Vidensk. Medd. Dansk Naturhist. Foren., 134: 191; and Wieczorek, Drewes, and Channing, 2001, Amphibia-Reptilia, 22: 159.

Rappia burgeoni De Witte, 1921, Rev. Zool. Afr., 9: 19. Holotype: MRAC 585, according to Laurent, 1943, Ann. Mus. R. Congo Belge, Tervuren, Zool., 4: 125. Type locality: "Madyu (Uelé)", Dem. Rep. Congo. Synonymy with Hyperolius viridiflavus pachyderma, by Laurent, 1951, Ann. Soc. R. Zool. Belg., 82: 23-50; Laurent, 1952 "1951", Ann. Soc. R. Zool. Belg., 82: 392; Inger, 1968, Explor. Parc Natl. Garamba, Miss. H. de Saeger, 52: 148. Dehling, 2015, Herpetozoa, Wien, 28: 39–47, noted that the types of Hyperolius pachydermus are conspecific with Hyperolius papyri, a member of the Hyperolius nasutus group, and explained how the name Rappia burgeoni remains a synonym of Hyperolius viridiflavus

Hyperolius burgeoniNoble, 1924, Bull. Am. Mus. Nat. Hist., 49: 252, 324; Ahl, 1931, Das Tierreich, 55: 314.

Hyperolius bayoniNoble, 1924, Bull. Am. Mus. Nat. Hist., 49: 252, 324.

Rappia rossii Calabresi, 1925, Atti Soc. Ital. Sci. Nat. Mus. Civ. Stor. Nat. Milano, 64: 121. Syntypes: MSNM 584 (2 specimens) according to Conci, 1967, Atti Soc. Ital. Sci. Nat. Mus. Civ. Stor. Nat. Milano, 106: 94; now MSNM Am 471 (2 specimens)  according to Blackburn and Scali, 2014, Herpetol. Monogr., 28: 30, who discussed the material. Type locality: "regione dell'alto Uellè (Congo Belga)" = Upper Uele Region, Dem. Rep. Congo. Corrected to "likely . . . from an area near or just east of present-day Watsa (3.037162 S, 29.535509 E)", Dem. Rep. Congo, by Blackburn and Scali, 2014, Herpetol. Monogr., 28: 30. Synonymy (with Hyperolius burgeoni) by Laurent, 1943, Ann. Mus. R. Congo Belge, Tervuren, Zool., 4: 67. Synonymy with Hyperolius marmoratus pachyderma by Laurent, 1951, Ann. Soc. R. Zool. Belg., 82: 23-50; Laurent, 1952 "1951", Ann. Soc. R. Zool. Belg., 82: 392. Dehling, 2015, Herpetozoa, Wien, 28: 39–47, noted that the types of Hyperolius pachydermus are conspecific with Hyperolius papyri, a member of the Hyperolius nasutus group, and explained how the name Rappia rossii remains a synonym of Hyperolius viridiflavus

Hyperolius rossiiAhl, 1931, Das Tierreich, 55: 283.

Hyperolius phrynoderma Ahl, 1931, Das Tierreich, 55: 344. Syntypes: ZMB (originally 5 specimens) unnumbered, according to the original publication; MCZ 17643 (on exchange from ZMB) is a syntype, according to Barbour and Loveridge, 1946, Bull. Mus. Comp. Zool., 96: 130). Type locality: "Zentrales Deutsch-Ostafrika". Placed in the synonymy of Hyperolius rossi by Loveridge, 1936, Bull. Mus. Comp. Zool., 79: 372. Considered a provisional synonym of Hyperolius marmoratus taeniatus by Laurent, 1961, Rev. Zool. Bot. Afr., 64: 83.The description in Ahl, 1931, Mitt. Zool. Mus. Berlin, 17: 71, appeared a few weeks later according to unpublished notes by A. Loveridge (fide R. Laurent).

Hyperolius schubotzi Ahl, 1931, Das Tierreich, 55: 328. Holotype: ZMB unnumbered, according to the original publication. Type locality: "Kissenji". Synonymy by Laurent, 1947, Bull. Mus. R. Hist. Nat. Belg., 23: 1-8; Laurent, 1983, Monit. Zool. Ital., N.S., Suppl., 18: 1-93. The description in Ahl, 1931, Mitt. Zool. Mus. Berlin, 17: 63, appeared a few weeks later according to unpublished notes by A. Loveridge (fide R. Laurent).

Hyperolius kwidjwiensis Ahl, 1931, Das Tierreich, 55: 296. Holotype: ZMB unnumbered according to the original publication. Type locality: "Insel Kwidjwi [=Kwidjwi Island] im Kivu-See [in Lake Kivu]", Dem. Rep. Congo. Synonymy with Hyperolius kivuensis by Loveridge, 1942, Bull. Mus. Comp. Zool., 91: 379, with Hyperolius viridiflavus schubotzi by Laurent, 1952 "1951", Ann. Soc. R. Zool. Belg., 82: 390; Laurent, 1983, Monit. Zool. Ital., N.S., Suppl., 18: 1-93. The description in Ahl, 1931, Mitt. Zool. Mus. Berlin, 17: 38, appeared a few weeks later according to unpublished notes by A. Loveridge (fide R. Laurent).

Hyperolius macrodactylus Ahl, 1931, Das Tierreich, 55: 368. Holotype: ZMB unnumbered, according to the original publication. Type locality: "Kivu-See". Synonymy with Hyperolius schubotzi by Loveridge, 1942, Bull. Mus. Comp. Zool., 91: 379. Synonymy with Hyperolius viridiflavus kwidjwiensis by Laurent, 1947, Bull. Mus. R. Hist. Nat. Belg., 23: 6. The description in Ahl, 1931, Mitt. Zool. Mus. Berlin, 17: 95, appeared a few weeks later according to unpublished notes by A. Loveridge (fide R. Laurent).

Hyperolius koehli Ahl, 1931, Das Tierreich, 55: 405. Holotype: ZMB unnumbered, according to the original publication; ZMB 26089, according to Laurent, 1961, Rev. Zool. Bot. Afr., 64: 91. Type locality: "Kissenji". Former synonymy (with Hyperolius kivuensis) by Loveridge, 1942, Bull. Mus. Comp. Zool., 91: 379. Synonymy (with Hyperolius viridiflavus schubotzi) by Laurent, 1950, Explor. Parc Natl. Albert, Miss. G.F. de Witte (1933–1935), 64: 93. The description in Ahl, 1931, Mitt. Zool. Mus. Berlin, 17: 121, appeared a few weeks later according to unpublished notes by A. Loveridge (fide R. Laurent).

Hyperolius variabilis Ahl, 1931, Das Tierreich, 55: 297. Holotype: ZMB unnumbered according to the original publication; MCZ 17648 (on exchange from ZMB) is a cotype according to Barbour and Loveridge, 1946, Bull. Mus. Comp. Zool., 96: 131. Type locality: "Deutsch-Ost-Afrika . . . Bukoba", Tanzania. Synonymy by Schiøtz, 1975, Treefrogs E. Afr.: 193. The description in Ahl, 1931, Mitt. Zool. Mus. Berlin, 17: 39, appeared a few weeks later according to unpublished notes by A. Loveridge (fide R. Laurent).

Hyperolius punctatissimus Ahl, 1931, Das Tierreich, 55: 299. Holotype: ZMB unnumbered, according to the original publication. Type locality: "Bukoba", Tanzania. Synonymy (with Hyperolius variabilis) by Laurent, 1943, Ann. Mus. R. Congo Belge, Tervuren, Zool., 4: 67. Synonymy with Hyperolius viridiflavus variabilis by Laurent, 1952 "1951", Ann. Soc. R. Zool. Belg., 82: 390; Laurent, 1961, Rev. Zool. Bot. Afr., 64: 90. (This disputed by Loveridge, 1957, Bull. Mus. Comp. Zool., 117: 330, who regarded it as a synonym of Hyperolius striolatus.) Synonymy by Schiøtz, 1975, Treefrogs E. Afr.: 41. The description in Ahl, 1931, Mitt. Zool. Mus. Berlin, 17: 41, appeared a few weeks later according to unpublished notes by A. Loveridge (fide R. Laurent).

Hyperolius wettsteini Ahl, 1931, Das Tierreich, 55: 343. Holotype: ZMB unnumbered, according to the original publication. Type locality: "Bukoba", Tanzania. Synonymy (with Hyperolius variabilis) by Laurent, 1943, Ann. Mus. R. Congo Belge, Tervuren, Zool., 4: 67. Synonymy with Hyperolius viridiflavus variabilis by Laurent, 1952 "1951", Ann. Soc. R. Zool. Belg., 82: 393; Laurent, 1961, Rev. Zool. Bot. Afr., 64: 90. Synonymy by Schiøtz, 1975, Treefrogs E. Afr.: 193. The description in Ahl, 1931, Mitt. Zool. Mus. Berlin, 17: 70, appeared a few weeks later according to unpublished notes by A. Loveridge (fide R. Laurent).

Hyperolius flavoguttatus Ahl, 1931, Das Tierreich, 55: 343. Holotype: ZMB unnumbered, according to the original publication; MCZ 17635 (on exchange from ZMB, according to Barbour and Loveridge, 1946, Bull. Mus. Comp. Zool., 96: 128) is presumably a paratype, not a syntype. Type locality: "Bukoba bis Kenia", Tanzania. Placed in the synonymy of Hyperolius viridiflavus variabilis by Laurent, 1952 "1951", Ann. Soc. R. Zool. Belg., 82: 393; Laurent, 1961, Rev. Zool. Bot. Afr., 64: 90. Synonymy by Schiøtz, 1975, Treefrogs E. Afr.: 193. The description in Ahl, 1931, Mitt. Zool. Mus. Berlin, 17: 96, appeared a few weeks later according to unpublished notes by A. Loveridge (fide R. Laurent).

Hyperolius irregularis Ahl, 1931, Das Tierreich, 55: 396. Syntypes: ZMB (2 specimens) unnumbered, according to the original publication. Type locality: "Mohasi-See, Ruanda". Synonymy by Schiøtz, 1975, Treefrogs E. Afr.: 193. Placed in the synonymy of Hyperolius rossi by Loveridge, 1936, Bull. Mus. Comp. Zool., 79: 372. Placed in the synonymy of Hyperolius variabilis) by Laurent, 1943, Ann. Mus. R. Congo Belge, Tervuren, Zool., 4: 67. Placed in the synonymy of Hyperolius viridiflavus variabilis by Laurent, 1952 "1951", Ann. Soc. R. Zool. Belg., 82: 393; Laurent, 1961, Rev. Zool. Bot. Afr., 64: 90. The description in Ahl, 1931, Mitt. Zool. Mus. Berlin, 17: 114, appeared a few weeks later according to unpublished notes by A. Loveridge (fide R. Laurent).

Hyperolius monticola Ahl, 1931, Das Tierreich, 55: 376. Holotype: ZMB unnumbered, according to the original publication. Type locality: "Niansa, Ruanda, 1500 m". Placed in the synonymy of Hyperolius variabilis) by Laurent, 1943, Ann. Mus. R. Congo Belge, Tervuren, Zool., 4: 67. In the synonymy of Hyperolius marmoratus variabilis according to Laurent, 1952 "1951", Ann. Soc. R. Zool. Belg., 82: 393. Synonymy by Schiøtz, 1975, Treefrogs E. Afr.: 193. The description in Ahl, 1931, Mitt. Zool. Mus. Berlin, 17: 102, appeared a few weeks later according to unpublished notes by A. Loveridge (fide R. Laurent).

Hyperolius mohasicus Ahl, 1931, Das Tierreich, 55: 360. Holotype: ZMB unnumbered, according to the original publication. Type locality: "Mohasi-See, Ruanda" (= Lake Mohasi, Rwanda). Synonymy by Schiøtz, 1975, Treefrogs E. Afr.: 193. Placed in the synonymy of Hyperolius variabilis) by Laurent, 1943, Ann. Mus. R. Congo Belge, Tervuren, Zool., 4: 67. Placed in the synonymy of Hyperolius rossi by Loveridge, 1936, Bull. Mus. Comp. Zool., 79: 372. In the synonymy of Hyperolius viridiflavus variabilis according to Laurent, 1952 "1951", Ann. Soc. R. Zool. Belg., 82: 393; Laurent, 1961, Rev. Zool. Bot. Afr., 64: 90. The description in Ahl, 1931, Mitt. Zool. Mus. Berlin, 17: 85, appeared a few weeks later according to unpublished notes by A. Loveridge (fide R. Laurent).

Hyperolius stuhlmanni Ahl, 1931, Das Tierreich, 55: 395. Holotype: ZMB unnumbered, according to the original publication and according to Laurent, 1961, Rev. Zool. Bot. Afr., 64: 91. Type locality: "Vitschumbi (Südende des Albert-Eduard-Sees)". Placed in synonymy of Hyperolius viridiflavus bayoni by Laurent, 1948 "1947", Ann. Mag. Nat. Hist., Ser. 11, 14: 288-294; Laurent, 1952 "1951", Ann. Soc. R. Zool. Belg., 82: 390; Laurent, 1961, Rev. Zool. Bot. Afr., 64: 91. Synonymy by Schiøtz, 1975, Treefrogs E. Afr.: 191. The description in Ahl, 1931, Mitt. Zool. Mus. Berlin, 17: 114, appeared a few weeks later according to unpublished notes by A. Loveridge (fide R. Laurent).

Hyperolius oculatus Ahl, 1931, Das Tierreich, 55: 377. Holotype: ZMB unnumbered, according to the original publication. Type locality: "Balaibo am Duki-Ufer". Placed in the synonymy of Hyperolius rossi by Loveridge, 1936, Bull. Mus. Comp. Zool., 79: 372. Placed in the synonymy of Hyperolius burgeoni by Laurent, 1943, Ann. Mus. R. Congo Belge, Tervuren, Zool., 4: 67. Considered a synonym of Hyperolius viridiflavus pachyderma by Laurent, 1952 "1951", Ann. Soc. R. Zool. Belg., 82: 390; Laurent, 1961, Rev. Zool. Bot. Afr., 64: 92. The description in Ahl, 1931, Mitt. Zool. Mus. Berlin, 17: 103, appeared a few weeks later according to unpublished notes by A. Loveridge (fide R. Laurent). Dehling, 2015, Herpetozoa, Wien, 28: 39–47, noted that the types of Hyperolius pachydermus are conspecific with Hyperolius papyri, a member of the Hyperolius nasutus group, and explained how the name Hyperolius oculatus remains a synonym of Hyperolius viridiflavus

Hyperolius karissimbiensis Ahl, 1931, Das Tierreich, 55: 348. Holotype: ZMB unnumbered, according to the original publication. Type locality: "Bambusurwald und Waldwiesen ca. 2400 m och, beim dorf des Mhcabu Gahama am Karissimbi", Kivu, Rwanda. Considered a subspecies of Hyperolius viridiflavus by Laurent, 1952 "1951", Ann. Soc. R. Zool. Belg., 82: 393; Laurent, 1983, Monit. Zool. Ital., N.S., Suppl., 18: 1-93. The description in Ahl, 1931, Mitt. Zool. Mus. Berlin, 17: 75, appeared a few weeks later according to unpublished notes by A. Loveridge (fide R. Laurent).

Hyperolius kandti Ahl, 1931, Das Tierreich, 55: 326. Holotype: ZMB unnumbered, according to the original publication. Type locality: "Kivu-See" (= Lake Kivu). Former synonymy with Hyperolius kivuensis by Loveridge, 1942, Bull. Mus. Comp. Zool., 91: 379. Synonymy with Hyperolius viridiflavus kwidjwiensis by Laurent, 1947, Bull. Mus. R. Hist. Nat. Belg., 23: 6. The description in Ahl, 1931, Mitt. Zool. Mus. Berlin, 17: 62, appeared a few weeks later according to unpublished notes by A. Loveridge (fide R. Laurent). ZMB

Hyperolius multicolor Ahl, 1931, Das Tierreich, 55: 367. Syntypes: ZMB (7 specimens) unnumbered, according to the original publication; MCZ 17641 (on exchange from ZMB, according to Barbour and Loveridge, 1946, Bull. Mus. Comp. Zool., 96: 129). Type locality: "Karissimbi, 2400 m". Synonymy (with Hyperolius karissimbiensis) by Laurent, 1950, Explor. Parc Natl. Albert, Miss. G.F. de Witte (1933–1935), 64: 76. Synonymy (with Hyperolius marmoratus karissimbiensis) by Laurent, 1952 "1951", Ann. Soc. R. Zool. Belg., 82: 394. The description in Ahl, 1931, Mitt. Zool. Mus. Berlin, 17: 94, appeared a few weeks later according to unpublished notes by A. Loveridge (fide R. Laurent).

Hyperolius taeniolatus Laurent, 1940, Rev. Zool. Bot. Afr., 34: 1-7. Holotype: MRAC 39471 by original designation. Type locality: "Lacs Mokoto", Dem. Rep. Congo. Preoccupied by Hyperolius taeniolatus Bocage, 1895. Synonymy (with Hyperolius viridiflavus coerulescens) by Laurent, 1950, Explor. Parc Natl. Albert, Miss. G.F. de Witte (1933–1935), 64: 96. Considered a synonym of Hyperolius viridiflavus by Laurent, 1947, Bull. Mus. R. Hist. Nat. Belg., 23: 8. Placed in the synonymy of Hyperolius variabilis) by Laurent, 1943, Ann. Mus. R. Congo Belge, Tervuren, Zool., 4: 67. Preoccupied by Rappia marmorata var. taeniolata Bocage, 1895 (= Hyperolius marmoratus taeniolatus of Laurent, 1952 "1951", Ann. Soc. R. Zool. Belg., 82: 390).

Hyperolius ornatus Laurent, 1940, Rev. Zool. Bot. Afr., 34: 4. Holotype: MRAC 3836, by original designation. Type locality: "Rutshuru", near Lake Kirwa, Kivu, Dem. Rep. Congo. Synonymy with Hyperolius schubotzi by Loveridge, 1942, Bull. Mus. Comp. Zool., 91: 379, 399. Synonymy (with Hyperolius viridiflavus bayoni) by Laurent, 1950, Explor. Parc Natl. Albert, Miss. G.F. de Witte (1933–1935), 64: 76; Laurent, 1952 "1951", Ann. Soc. R. Zool. Belg., 82: 390. Synonymy with Hyperolius viridiflavus by Laurent, 1972, Explor. Parc Natl. Virunga, Ser. 2, 22: 84.

Hyperolius schubotzi ruandensis Laurent, 1943, Ann. Mus. R. Congo Belge, Tervuren, Zool., 4: 129. Holotype: MRAC 38371, by original designation. Type locality: "Nyakatare (Ruanda [= Rwanda])". Synonymy with Hyperolius viridiflavus variabilis by Laurent, 1983, Monit. Zool. Ital., N.S., Suppl., 18: 1-93.

Hyperolius ornatus taeniolatusLaurent, 1943, Ann. Mus. R. Congo Belge, Tervuren, Zool., 4: 131.

Hyperolius destefanii Scortecci, 1943, in Zavattari (ed.), Miss. Biol. Sagan–Omo, 7(Zool. 1): 321. Holotype: MSNM, but likely now lost, according to Conci, 1967, Atti Soc. Ital. Sci. Nat. Mus. Civ. Stor. Nat. Milano, 106: 94, and Blackburn and Scali, 2014, Herpetol. Monogr., 28: 37. Type locality: "Nargi", 04° 48′ N, 36° 08′ E, Ethiopia. A member of the Hyperolius viridiflavus superspecies according to Schiøtz, 1999, Treefrogs Afr.: 208.

Hyperolius viridiflavus coerulescens Laurent, 1947, Bull. Mus. R. Hist. Nat. Belg., 23: 8. Replacement name for Hyperolius taeniolatus Laurent, 1940.

Hyperolius viridiflavus kwidjwiensisLaurent, 1947, Bull. Mus. R. Hist. Nat. Belg., 23: 6.

Hyperolius viridiflavus viridiflavusLaurent, 1947, Bull. Mus. R. Hist. Nat. Belg., 23: 8; Laurent, 1951, Ann. Soc. R. Zool. Belg., 82: 40; Loveridge, 1957, Bull. Mus. Comp. Zool., 117: 324; Schiøtz, 1975, Treefrogs E. Afr.: 188.

Hyperolius viridiflavus bayoniLaurent, 1947, Bull. Mus. R. Hist. Nat. Belg., 23: 8; Laurent, 1950, Explor. Parc Natl. Albert, Miss. G.F. de Witte (1933–1935), 64: 85.

Hyperolius viridiflavus schubotziLaurent, 1947, Bull. Mus. R. Hist. Nat. Belg., 23: 1–8; Laurent, 1951, Ann. Soc. R. Zool. Belg., 82: 41; Laurent, 1983, Monit. Zool. Ital., N.S., Suppl., 18: 1-93.

Hyperolius karissimbiensis xanthogrammus Laurent, 1950, Rev. Zool. Bot. Afr., 44: 9. Holotype: MRAC unnumbered by original designation. Type locality: "Cyumba, Marais de la Mulindi (Alt. 1. 950 m.), Territoire de Biumba, Ruanda [= Rwanda]". Transfer to subspecies of Hyperolius viridiflavus by Laurent, 1952 "1951", Ann. Soc. R. Zool. Belg., 82: 394.

Hyperolius karissimbiensis françoisi Laurent, 1950, Rev. Zool. Bot. Afr., 44: 10. Holotype: MRAC unnumbered by original designation. Type locality: "Bururi (Alt. 1.850-1.950 m.), Urundi". Transfer to subspecies of Hyperolius viridiflavus by Laurent, 1952 "1951", Ann. Soc. R. Zool. Belg., 82: 394.

Hyperolius karissimbiensis intermedius Laurent, 1950, Rev. Zool. Bot. Afr., 44: 11. Holotype: MRAC unnumbered by original designation. Type locality: "Marais Rugushu, Kayanza (Alt. 1.900 m.), Territoire de Ngozi, Urundi [Burundi]". Transferred to subspecies of Hyperolius viridiflavus by Laurent, 1952 "1951", Ann. Soc. R. Zool. Belg., 82: 394. Considered a junior synonym of Hyperolius viridiflavus francoisi by Laurent, 1983, Monit. Zool. Ital., N.S., Suppl., 18: 1-93.

Hyperolius viridiflavus hybridus Laurent, 1951, Rev. Zool. Bot. Afr., 44: 375. Holotype: MRAC unnumbered, according to the original publication. Type locality: "Estand de la Mukura, 1.700 m., près d'Astrida, Ruanda".

Hyperolius viridiflavus variabilisLaurent, 1951, Ann. Soc. R. Zool. Belg., 82: 41.

Hyperolius viridiflavus xanthogrammusLaurent, 1951, Ann. Soc. R. Zool. Belg., 82: 41; Laurent, 1972, Explor. Parc Natl. Virunga, Ser. 2, 22: 85.

Hyperolius viridiflavus angeli Laurent, 1951, Ann. Soc. R. Zool. Belg., 82: 41. Holotype: MNHNP 1921.34b, by original designation. Type locality: "Gribingui, Oubangi-Chari".

Hyperolius viridiflavus variabilisLaurent, 1951, Ann. Soc. R. Zool. Belg., 82: 41; Laurent, 1983, Monit. Zool. Ital., N.S., Suppl., 18: 1-93.

Hyperolius kandiLaurent, 1952 "1951", Ann. Soc. R. Zool. Belg., 82: 393. Typographic error.

Hyperolius viridiflavus xanthogrammusLaurent, 1952 "1951", Ann. Soc. R. Zool. Belg., 82: 394.

Hyperolius viridiflavus françoisiLaurent, 1952 "1951", Ann. Soc. R. Zool. Belg., 82: 394.

Hyperolius viridiflavus intermediusLaurent, 1952 "1951", Ann. Soc. R. Zool. Belg., 82: 394.

Hyperolius bayoniLoveridge, 1957, Bull. Mus. Comp. Zool., 117: 326.

Hyperolius viridiflavus ornatusLaurent, 1972, Explor. Parc Natl. Virunga, Ser. 2, 22: 4, 84.

Hyperolius viridiflavus mwanzae Schiøtz, 1975, Treefrogs E. Afr.: 194. Holotype: ZMUC R078836, by original designation. Type locality: "swamp north of Mwanza, Tanzania".

Hyperolius viridiflavus karissimbiensisLaurent, 1983, Monit. Zool. Ital., N.S., Suppl., 18: 1-93.

Hyperolius viridiflavus francoisiLaurent, 1983, Monit. Zool. Ital., N.S., Suppl., 18: 1-93.

Hyperolius mwanzaePickersgill, 2007, Frog Search: 418; Schiøtz, 1999, Treefrogs Afr.: 42-46.

English Names

Common Reed Frog (Hyperolius viridiflavus: Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 68; Channing and Howell, 2006, Amph. E. Afr.: 186).

Common Tree Frog (Hyperolius viridiflavus: Lötters, Wagner, Bwong, Schick, Malonza, Muchai, Wasonga, and Veith, 2007, Fieldguide Amph. Rept. Kakamega Forest: 34).

Kenyan Reed Frog (Hyperolius viridiflavus: Ananjeva, Borkin, Darevsky, and Orlov, 1988, Dict. Amph. Rept. Five Languages: 74).

Tigoni Reed Frog (Hyperolius destefani [no longer recognized]: Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 67).

Distribution

East Africa in eastern Dem. Rep. Congo west to north-east-central Central African Republic, Uganda, Tanzania, western Ethiopia almost to the Eritrea border, and southern Somalia, South Sudan, into extreme southern Sudan.

Comment

In the Hyperolius viridiflavus superspecies according to Schiøtz, 1975, Treefrogs E. Afr.: 172. See also Schiøtz, 1971, Vidensk. Medd. Dansk Naturhist. Foren., 134: 21-76; and Laurent, 1983, Monit. Zool. Ital., N.S., Suppl., 18: 1–93. See general discussion of the Hyperolius viridiflavus–parallelus–marginatus group by Schiøtz, 1999, Treefrogs Afr.: 206–228, and specifically (including the forms he calls Pachydermus, Viridiflavus, Bayoni, Destefanii, Variabilis, Coerulescens, Karissimbiensis, Françoisi, and Mwanzae), although this has been modified by subsequent authors; see particular Hyperolius glandicolor. See comments under Hyperolius marmoratus, and Hyperolius sheldricki. See account of Ethiopian population by Largen, 1998, Tropical Zool., 11: 61-82. Wieczorek, Drewes, and Channing, 2000, J. Biogeograph., 27: 1241, provided a map and a considerable altered concept of this species. Lanza, 1990, Biogeographia, 14: 411, noted the occurrence of this species in southern Somalia. Channing and Howell, 2006, Amph. E. Afr.: 186, and Lötters, Wagner, Bwong, Schick, Malonza, Muchai, Wasonga, and Veith, 2007, Fieldguide Amph. Rept. Kakamega Forest: 34-36, provided accounts. See account, photograph, and map for Ethiopia by Largen and Spawls, 2010, Amph. Rept. Ethiopia Eritrea: 118-120. Channing, Rödel, and Channing, 2012, Tadpoles of Africa: 227, provided information on comparative larval morphology.

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