Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Heterixalus variabilis (Ahl, 1930)

Class: Amphibia > Order: Anura > Family: Hyperoliidae > Subfamily: Hyperoliinae > Genus: Heterixalus > Species: Heterixalus variabilis

Megalixalus variabilis Ahl, 1930, Mitt. Zool. Mus. Berlin, 16: 526. Syntypes: ZMB (12 specimens), unnumbered in the original publication; including ZMB 7856, 50108-117, a total 11 specimens, according to Tillack, de Ruiter, and Rödel, 2021, Zoosyst. Evol., Berlin, 97: 440. Type locality: "Nossi Faly [Nosy Faly, SW Ambaro Bay, Ambanja District, Diana Region], bei Madagascar".

Heterixalus variabilis — Laurent, 1950, Explor. Parc Natl. Albert, Miss. G.F. de Witte (1933–1935), 64: 14, by implication; Bachmann and Blommers-Schlösser, 1975, Zool. Anz., 194: 21; based on specimens of Heterixalus alboguttatus, according to Blommers-Schlösser, 1982, Beaufortia, 32: 9; Glaw and Vences, 1992, Fieldguide Amph. Rept. Madagascar: 278.

English Names

None noted.

Distribution

Ambanja and Ambilobe, Madagascar.

Geographic Occurrence

Natural Resident: Madagascar

Endemic: Madagascar

Comment

See Blommers-Schlösser and Blanc, 1991, Faune de Madagascar, 75: 126. Formerly included in the synonymy of Heterixalus alboguttatus by Blommers-Schlösser, 1982, Beaufortia, 32: 9, and considered a nomen dubium by Guibé, 1978, Bonn. Zool. Monogr., 11: 87, although Glaw and Vences, 1993, Salamandra, 29: 212-230, applied this name to this species. F. Glaw (pers. comm.) suggests that the validity of this name is still uncertain and may be a synonym of Heterixalus tricolor. Glaw and Vences, 1994, Fieldguide Amph. Rept. Madagascar, Ed. 2: 77, discussed the problems with application of this name. Glaw and Vences, 2007, Field Guide Amph. Rept. Madagascar, Ed. 3: 104, provided an account. Tillack, de Ruiter, and Rödel, 2021, Zoosyst. Evol., Berlin, 97: 440, briefly discussed the syntypes and their collector.

External links:

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For access to general information see Wikipedia
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For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.