Hyla intermedia Boulenger, 1882

Class: Amphibia > Order: Anura > Family: Hylidae > Subfamily: Hylinae > Genus: Hyla > Species: Hyla intermedia

Hylaria variegata Rafinesque, 1814, Specchio Sci., 2, 2: 103. Types: Not designated and not known to exist. Neotype, by subsequent designation of Nascetti, Lanza, and Bullini, 1995, Amphibia-Reptilia, 16: 225, MZUF 20365. Type locality: "comune nei giardini vicino Palermo e Monreale", Sicily, Italy. Neotype from "the wood called Bosco della Ficuzza, about 10 km by air NE of Corleone (Palermo; ca. 37° 00′ N - 14° 20′ E), altitude about 1000 m". Secondary homonym of Rana variegata Bonnaterre, 1789 (= Hyla leucophyllata).

Hyla arborea var. intermedia Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 381. Syntypes: BMNH (Bononia), (Palermo); BMNH designated lectotype by Dubois, 1995, Dumerilia, 2: 67. Type locality: "Bononia" and "Palermo, Sicily", Italy; restricted to "Palermo" by lectotype designation. Synonymy by Dubois, 1995, Dumerilia, 2: 55-71.

Hyla intermediaHerón-Royer, 1890, Bull. Soc. Zool. France, 15: 209.

Hyla italica Nascetti, Lanza, and Bullini, 1995, Amphibia-Reptilia, 16: 225. Replacement name for Hylaria variegata Rafinesque, 1814. Synonymy with Hyla intermedia by Dubois, 1995, Dumerilia, 2: 55-71.

Hyla (Hyla) intermedia — Fouquette and Dubois, 2014, Checklist N.A. Amph. Rept.: 331, by implication. 

English Names

Italian Tree Frog (Arnold, 2002, Rept. Amph. Eur., Ed. 2: 78).


Italy south of the Po Valley south into Sicily. 


In the Hyla arborea group of Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005, Bull. Am. Mus. Nat. Hist., 294: 102 (see comment under Hyla). See Dubois, 1995, Dumerilia, 2: 55-71, for extended discussion of nomenclature. Lanza, 1983, Guide Reconoscimento Spec. Animali, Anf. Rett.: 110-113, discussed this species as Hyla arborea. Arnold, 2002, Rept. Amph. Eur., Ed. 2: 74-75, provided a brief account, figure, and map. Canestrelli, Cimmaruta, and Nascetti, 2007, Mol. Ecol., 16: 4808-4821, reported on molecular phylogeography. Stöck, Dubey, Klütsch, Litvinchuk, Scheidt, and Perrin, 2008, Mol. Phylogenet. Evol., 49: 1019-1024, reported on molecular phylogenetics and noted that it is composed of two distinctive subclades. See detailed account for Italy by Lapini, 2007, in Lanza et al. (eds.), Fauna d'Italia, 42 (Amph.): 333-338. See comments by Speybroeck, Beukema, and Crochet, 2010, Zootaxa, 2492: 7, on the evidence of cryptic taxa within this nominal species. Grosse, 2011, Laubfrösche: 1-286, reviewed the biology of the species. Dufresnes, Dubey, Ghali, Canestrelli, and Perrin, 2015, Conserv. Genetics, 16: 1507–1513, reported on introgressive hybridization of an introduced populations with Hyla arborea in western Switzerland.  Speybroeck, Beukema, Bok, and Van Der Voort, 2016, Field Guide Amph. Rept. Brit. Eur.: 158–159, provided a brief account and range map. Speybroeck, Beukema, Bok, and Van Der Voort, 2016, Field Guide Amph. Rept. Brit. Eur.: 162–164, provided a brief account and range map. Dufresnes, Mazepa, Rodrigues, Brelsford, Litvinchuk, Sermier, Lavanchy, Betto-Colliard, Blaser, Borzée, Cavoto, Fabre, Ghali, Grossen, Horn, Leuenberger, Phillips, Saunders, Savary, Maddalena, Stöck, Dubey, Canestrelli, and Jeffries, 2018, Frontiers Ecol. Evol., 6 (144): 1–18, reported on the molecular phylogeography of the species, resulting in the northern (Po Valley) population being recognized as Hyla perrini, with which it hybridizes on the northeastern edge of the Apennines in Emilia Romagna. 

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