Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Craugastor rugulosus (Cope, 1870)

Class: Amphibia > Order: Anura > Superfamily: Brachycephaloidea > Family: Craugastoridae > Genus: Craugastor > Species: Craugastor rugulosus

Liyla rugulosa Cope, 1870 "1869", Proc. Am. Philos. Soc., 11: 160. Syntypes: USNM 29971-72 according to Kellogg, 1932, Bull. U.S. Natl. Mus., 160: 117; USNM 29771 designated lectotype by Savage, 1974, Herpetologica, 30: 295, and Savage, 1975, Copeia, 1975: 273. Type locality: "Pacific region of the isthmus of Tehuantepec", Mexico.

Hylodes palmatus Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 201. Syntypes: By indication (as a composite replacement name) the primary types of Leihyla güntheri Keferstein (= Eleutherodactylus fitzingeri) and Liyla rugulosa Cope (= Eleutherodactylus rugulosus). Among the types of these species USNM 29771 was designated lectotype of Hylodes palmatus by Savage, 1974, Herpetologica, 30: 295. Type locality: "Pacific region of the Isthmus of Tehuantepec", Mexico. Synonymy by Savage, 1974, Herpetologica, 30: 295.

Eleutherodactylus palmatus — Stejneger, 1904, Annu. Rep. U.S. Natl. Mus. for 1902: 582-583, by implication; Taylor, 1952, Univ. Kansas Sci. Bull., 35: 744 (Based on specimens of Eleutherodactylus andi, according to Savage, 1974, Herpetologica, 30: 295).

Eleutherodactylus rugulosus — Kellogg, 1932, Bull. U.S. Natl. Mus., 160: 116.

Eleutherodactylus avocalis Taylor and Smith, 1945, Proc. U.S. Natl. Mus., 95: 580. Holotype: USNM 116885. Type locality: "Tres Cruces, near Tehuantepec, Oaxaca", Mexico. Synonymy by Lynch, 1965, Herpetologica, 21: 110, and Savage, 1975, Copeia, 1975: 273.

Eleutherodactylus rugulosus rugulosus — Duellman, 1961, Univ. Kansas Publ. Mus. Nat. Hist., 15: 38-39, by implication.

Eleutherodactylus (Craugastor) rugulosus — Hedges, 1989, in Woods (ed.), Biogeograph. W. Indies: 317, by implication; Lynch, 1996, in Powell and Henderson (eds.), Contr. W. Indian Herpetol.: 154.

Craugastor rugulosus — Crawford and Smith, 2005, Mol. Phylogenet. Evol., 35: 551, by implication; Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 360.

Craugastor (Craugastor) regulosus — Hedges, Duellman, and Heinicke, 2008, Zootaxa, 1737: 41.

English Names

Central American Rain Frog (Liner, 1994, Herpetol. Circ., 23: 21; Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 79).

Rugulose Rainfrog (Lee, 1996, Amph. Rept. Yucatan Peninsula: 65; Liner and Casas-Andreu, 2008, Herpetol. Circ., 38: 11).

Distribution

Slopes of the Pacific drainage of southern Mexico from central Guerrero, where it occurs in the Río Balsas and Río Papagayo drainages, eastward through Morelos, southern Puebla and southern Oaxaca into extreme southwestern Chiapas, 200-over 2000 m elevation.

Geographic Occurrence

Natural Resident: Mexico

Endemic: Mexico

Comment

See comment under Eleutherodactylus berkenbuschii. See Campbell and Savage, 2000, Herpetol. Monogr., 14: 204-208, for synonymy and review. See comment under Eleutherodactylus brocchi. In the Eleutherodactylus (Craugastor) rugulosus group, according to Lynch and Duellman, 1997, Univ. Kansas Mus. Nat. Hist. Spec. Publ., 23: 232. Savage, 2002, Amph. Rept. Costa Rica: 226, included this species in his Eleutherodactylus fitzingeri series, Eleutherodactylus rugulosus group. Valenzuela Galván, Chávez-Juárez, and Alcaraz-Cruz, 2004, Herpetol. Rev., 35: 184, provided the Morelos record. In the Craugastor punctariolus species series of Hedges, Duellman, and Heinicke, 2008, Zootaxa, 1737: 42, and of Padial, Grant, and Frost, 2014, Zootaxa, 3825: 123.

External links:

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For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.