Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Cardioglossa gracilis Boulenger, 1900

Class: Amphibia > Order: Anura > Family: Arthroleptidae > Subfamily: Arthroleptinae > Genus: Cardioglossa > Species: Cardioglossa gracilis

Cardioglossa gracilis Boulenger, 1900, Proc. Zool. Soc. London, 1900: 446. Syntypes: BMNH 1947.2.30.39–43 (formerly 1900.2.17.77–81), IRSNB 1.717 (formerly BMNH 1947.2.17.82), according to Amiet, 1985, in Frost (ed.), Amph. Species World: 16, and Lang, 1990, Doc. Trav., Inst. R. Sci. Nat. Belg., 59: 6. Type locality: "Benito River", elsewhere in the original publication noted as "Benito River, north of the Gaboon River between 20 and 30 miles inland from the coast, Gaboon" (actually in Equatorial Guinea).

Astylosternus gracilis — Dubois, Ohler, and Pyron, 2021, Megataxa, 5: 222, by implication. See comment under Cardioglossa.

English Names

Rio Benito Long-fingered Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 37).

Gracile Long-fingered Frog (Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 240).

Distribution

Southeastern Nigeria to northwestern Dem. Rep. Congo (and in adjacent southwestern Central African Republic) through southern Cameroon, Equatorial Guinea, and Gabon, in forested areas under 1200 m elevation; reported from Uigé Province of northwestern Angola; presumably to be found in intervening Rep. Congo

Geographic Occurrence

Natural Resident: Angola, Cameroon, Central African Republic, Congo, Democratic Republic of the, Equatorial Guinea, Gabon, Nigeria

Likely/Controversially Present: Congo, Republic of the

Comment

Frétey and Blanc, 2002 "2001", Bull. Soc. Zool. France, 126: 381, reported this species from Gabon. Lasso, Rial, Castroviejo, and De la Riva, 2002, Graellsia, 58: 21–34, provided notes on ecological distribution in Equatorial Guinea. Schiøtz, 1963, Vidensk. Medd. Dansk Naturhist. Foren., 125: 32–33, provided records for Nigeria. Jongsma, Tobi, Dixon-MacCallum, Bamba-Kaya, Yoga, Mbega, Mve Beh, Emrich, and Blackburn, 2017, Amph. Rept. Conserv., 11 (1: e144): 6, provided a record for southeastern Gabon and briefly discussed habitat and range. Larson and Zimkus, 2018, Herpetol. Conserv. Biol., 13: 246, reported a specimen from Birougou, Gabon. Dewynter, Chirio, Melki, Cordier, and Frétey, 2017, Cah. Fondation Biotope, 11: 10, provided a brief account and photographs of this species from the Mount Koumouna-Bouali area of Gabon. Dewynter, Frétey, Jongsma, Bamba-Kaya, and Pauwels, 2018, Cah. Fondation Biotope, 18: 23, provided a brief account and photographs from the Monts Birougou area of Gabon. Dewynter and Frétey, 2019, Cah. Fondation Biotope, 27: 9, summarized the literature for Gabon and provided a photograph. Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 240–241, provided a brief account, photograph, and range map. See brief account, range map, and photographs for Equatorial Guinea in Sánchez-Vialas, Calvo-Revuelta, Castroviejo-Fisher, and De la Riva, 2020, Proc. California Acad. Sci., Ser. 4, 66: 137–230. Ernst, Lautenschläger, Branquima, and Hölting, 2020, Zoosyst. Evol., 96: 249, noted the species to occur in Uigé Province of northwestern Angola.

External links:

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For access to general information see Wikipedia
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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.