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Tylototriton asperrimus Unterstein, 1930

Class: Amphibia > Order: Caudata > Family: Salamandridae > Subfamily: Pleurodelinae > Genus: Tylototriton

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Tylototriton asperrimus Unterstein, 1930, Sitzungsber. Ges. Naturforsch. Freunde Berlin, 1930: 314. Syntypes: ZMB (2 specimens); ZMB 34089 considered holotype (lectotype designation by implication) by Bauer, Good, and Günther, 1993, Mitt. Zool. Mus. Berlin, 69: 298. Type locality: "Kwangsi"; corrected to "Yao Shan [= Mt. Yao], Kwangsi Prov. [= Guangxi Zhuang Autonomous Region], China" by Bauer, Good, and Günther, 1993, Mitt. Zool. Mus. Berlin, 69: 298. Fan, 1931, Bull. Dept. Biol. Coll. Sci. Sun Yatsen Univ., 11: 1-154, provided evidence that the type locality is or near to the town of Loshiang.

Tylototriton (Echinotriton) asperrimus — Zhao and Hu, 1984, Stud. Chinese Tailed Amph.: 19.

Tylototriton asperrimus asperrimus — Fei, Ye, and Yang, 1984, Acta Zool. Sinica, 30: 89.

Pleurodeles (Tylototrion) asperrimus asperrimus — Risch, 1985, J. Bengal Nat. Hist. Soc., N.S.,, 4: 142.

Pleurodeles (Echinotriton) asperrimus — Dubois, 1987 "1986", Alytes, 5: 11.

Echinotriton asperrimus — Zhao, 1990, in Zhao (ed.), From Water onto Land: 219. Nguyen, Ho, and Nguyen, 2005, Checklist Amph. Rept. Vietnam: 9.

Echinotriton asperrimus asperrimus — Zhao and Adler, 1993, Herpetol. China: 112.

Tylototriton asperrimus — Nussbaum, Brodie, and Yang, 1995, Herpetologica, 51: 265.

Tylototriton (Yaotriton) asperrimus — Dubois and Raffaëlli, 2009, Alytes, 26: 68.

English Names

Mount Yao Newt (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 35).

Black Knobby Newt (Fei, 1999, Atlas Amph. China: 40).

Granular Newt (Nguyen, Ho, and Nguyen, 2005, Checklist Amph. Rept. Vietnam: 9).

Distribution

Manson Mountains of Anhui, Guangxi, Guangdong, Gansu,Hunan, Sichuan, and Hubei, China; Mau Son and Lang Son areas of Ha Giang and Laco Cai provinces in northern Vietnam, 400-1700 m elevation.

Comment

See accounts by Bourret, 1942, Batr. Indochine: 146-148; Fang and Chang, 1932, Sinensia, Nanking, 2: 117-121; Ye, Fei, and Hu, 1993, Rare and Economic Amph. China 72; and Fei, 1999, Atlas Amph. China: 40. See also Liu, Hu, Fei, and Huang, 1973, Acta Zool. Sinica, 19: 394. Deng and Yu, 1984, Acta Herpetol. Sinica, Chengdu, N.S.,, 3 (2): 75-77, recognize a subspecies in Sichuan. Nussbaum, Brodie, and Yang, 1995, Herpetologica, 51: 265, discussed reasons for transferring this species from Echinotriton to Tylototriton. Thorn and Raffaëlli, 2000, Salamand. Ancien Monde: 236-238, provided an account (as Echinotriton asperrimus asperrimus). Zhang and Wen, 2000, Amph. Guangxi: 21, provided an account for population in Guangxi, China. Bain and Nguyen, 2004, Am. Mus. Novit., 3453: 9, provided the record for Ha Giang Province, Vietnam. Nguyen, Ho, and Nguyen, 2005, Checklist Amph. Rept. Vietnam: 9, provided specific records for Vietnam and (p. 147) provided a photograph. In the Tylototriton asperrimus group of Fei, Ye, Huang, Jiang, and Xie, 2005, Illust. Key Chinese Amph.: 42 (although they only addressed Chinese species). Böhme, Schöttler, Nguyen, and Köhler, 2005, Salamandra, 41:218-219, discussed previous confusion in Vietnam of Tylototriton asperrimus and Tylototriton vietnamensis and noted that the record for Tam Dao, northern Vietnam, may actually be referrable to Tylototriton hainanensis. Fei, Hu, Ye, and Huang, 2006, Fauna Sinica, Amph. 1: 257-261, provided an account. Raffaëlli, 2007, Les Urodèles du Monde: 147-148, provided a brief account, figure, and map. See statement of geographic range, habitat, and conservation status in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 643. Nguyn, Nguyn, H, Lê, and Nguyn, 2009, Tp chí Công ngh Sinh hc, 7: 325-333, discussed the species in Vietnam. Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 72-73, provided a brief account including photographs of specimens and habitat.

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